Rozprawy doktorskie na temat „Radiation interception”

The potential of agroforestry to alleviate problems related to scarcities of arable land, water, food and fuel wood is subject to understanding system functioning and implementing and managing an efficiently designed system. The objectives of this study were to understand interactions and productivity of a hedgerow intercropping system with reference to water and radiation use, and analyse system design and management scenarios in order to enhance returns. Field trials monitoring soil water, solar radiation and plant productivity were conducted during 2006-2008 at Ukulinga Research Farm (KwaZulu Natal, South Africa) using a Jatropha-Kikuyu (Pennisetum clandestinum) hedgerow intercropping system as case study. In order to extrapolate results, a process-based hedgerow intercropping model was developed by building intercropping and tree growth into the SWB-2D model. Data collected from the field trials were used to parameterise and evaluate the model, which was used to analyse hedgerow orientation and spacing to determine income scenarios of virtual system and to help develop design criteria. Allometric relationships of Jatropha using basal stem diameter and crown width as predictor variables were found to be very reliable. Stem diameter was linearly related with wood and branch proportions and inversely proportional to foliage. Neither below-ground (BG) interspecies competition nor tree spacing had any significant effects on allometry. Allometric equations were proven valid for accurate, non-destructive and rapid predictions of tree growth under various growing and non-destructive canopy management conditions. When interspecies competition was present, none of the tree spacing/arrangement options tested resulted in consistently highest tree relative growth rates (RGR). Treatments had no effect on tree RGR when high water availability and kikuyu dormancy coincided. The single-row treatment (SR) produced the shortest trees, but generally had the highest stem RGR during low rainfall periods. The standard-spacing treatment (SS) had the highest RGR during the spring and summer seasons. Jatropha-only treatment (JO) trees were the tallest and biggest. Treatments affected post-pruning tree height increase, even when rainfall was high. Length of tree-crop interface (TCI) generally decreased tree yield, especially as trees matured toward their maximum-yield age (4-5 years). SR trees showed slow response to pruning due to a high TCI. They, however, exhibited compensatory growth during May to August, when competition for water with grass was low. BG competition reduced tree nut yield more than tree biomass. Tree spacing/arrangements had no effect on tree harvest index. Soil water varied among treatments and was asymmetrically distributed across tree hedgerows. System ET was generally the highest in SR and lowest in the double-row treatment (DR). Differences were mainly due to transpiration. Treatments affected tree root distribution, which was inferred using correlations between tree RGR and soil water deficit (SWD). In JO and SR, fine tree roots were asymmetrically distributed. Their distribution in DR was essentially symmetrical. Strong vegetative RGR-SWD correlations during the 2007/08 season indicated that tree growth was mainly water-limited. Though DR and SR had comparable tree RGRs, DR produced less grass than SR. This implied DR had more intensive BG competition than SR. Interspecific competition was severe due to a lack of temporal complementarity between Jatropha and kikuyu and a shallow soil profile (0.6 m). Tree water uptake predominantly came from the 0.2 – 0.6 depth, which had about 8.6% of the total root biomass in the profile. There was no clear relationship between intercrop growth and root distribution. Radiation use efficiency of kikuyu decreased towards tree hedgerows possibly due to preceding interaction of the irradiance with tree canopy reducing photosynthetically active radiation. The effect of radiation distribution on tree-crop (T-C) interactions was mainly to magnify effects of water. Finally, tree spacing/arrangement could be manipulated to optimise radiation and soil water distribution and intercrop growth. Predictions of solar radiation distribution, profile water content and tree water use were quite accurate. In general, intercrop productivity simulations were acceptable. Intercrop growth was overestimated when rainfall was high and underestimated when rainfall was low. During model calibration, tree woody biomass, leaf area index, crown width and nut yield were predicted adequately, while leaf dry mass was overestimated. During model validation, woody biomass and crown width were simulated reasonably well. However, foliage biomass, leaf area index and nut yield were overestimated. Overall, adequacy of the model for simulating tree productivity was established. Using scenario modelling, model capabilities to facilitate design/planning and management of hedgerow intercropping systems and interpretation of model outputs were demonstrated. The model can be used to determine the T-C trade-off that yields maximum income. By selecting best-case row orientation and spacing scenarios using the model, and keeping in mind values of tree and intercrop yields, system returns can be maximised. Tree crown growth can also be predicted in order to decide on the extent and timing of pruning. The present model is applicable to any potential tree-intercrop combination. It should be linked to a nutrient simulator of SWB, its component, and appraised further by considering shade-intolerant and shade-loving crop species, along with evergreen and deciduous tree species. This provides model users with numerous T-C combinations to choose from. Various tree spacing/arrangement options can also be explored using the model in order to realise the full potential and implications of the experimental findings of this study and others.
Thesis (PhD)--University of Pretoria, 2012.
gm2014
Plant Production and Soil Science
unrestricted

Les surchauffes urbaines,additionnées aux vagues de chaleur dont la fréquence et l’intensité augmentent en raison du changement climatique, exposent les citadins à un stress thermique croissant. Les arbres d’alignement peuvent permettre d’atténuer ces surchauffes grâce à l’ombrage et la transpiration,deux phénomènes qui peuvent dépendre de leurs caractéristiques morphologiques et physiologiques. En parallèle, les périodes de sécheresse se multiplient et la capacité des arbres d’alignement à croître tout en maintenant des services climatiques importants dans des conditions chaudes et sèches est en jeu. Pour améliorer la compréhension des facteurs influant sur les bénéfices climatiques des arbres, une démarche de recherche basée sur la mesure des interactions le long du continuum sol-arbre atmosphère dans deux dispositifs expérimentaux complémentaires est proposée. D’un côté, une rue canyon à échelle 1/5èmepartiellement végétalisée avec des jeunes pommiers d’ornement (Malus Coccinella® ‘Courtarou’) en pot et irrigués située à Angers(France) a été instrumentée pour quantifier les effets sur le microclimat d’arbres soumis à différentes disponibilités en eau. De l’autre, une campagne de mesures in-situ du microclimat dans et autour de la couronne de 36 arbres d’alignement adultes de 9 essences différentes répartis sur 5 sites dans Paris (France) a été menée. L’analyse croisée de ces deux dispositifs fournit une vue d’ensemble des sources de variabilité des bénéfices climatiques et propose aux aménageurs une meilleure compréhension des liens entre la résistance à la sécheresse des arbres d’alignement et leur pouvoir de réduction du stress thermiquehumain
Cities are characterized by an urban overheating which, combined to more frequent and intense heatwaves caused by climate change, expose the city dwellers to an increasing human thermal stress. Street trees may partially mitigate overheating by means of cast shadows and transpiration, two processes which may depend on their morphological and physiological characteristics. At the same time, periods of severe water shortage are multiplying and the ability of street trees to thrive and maintain significant cooling benefits under heat and drought conditions is crucial. In order to improve the understanding of the factors influencing tree climatic benefits, a methodology based on the monitoring of the interactions along the soil-tree atmospherecontinuum in two complementary experimental facilities was developed. On the one hand, a 1/5 scale canyon street, partially planted with young potted drip-irrigated ornamental apple trees (Malus Coccinella® ‘Courtarou’) located in Angers, France was monitored to quantify the effect of street trees experiencing different water supplies on the microclimate. On the other hand, an in-situ monitoring of the microclimate inside and outside the canopy for 36 mature street trees from 9 different tree species located in 5 sites in Paris, France, was performed. On the basis of a detailed cross-analysis of these two experimental facilities, an integrated assessment of the street trees climatic benefits was proposed to explore the sources of variability of the climatic benefits provided by trees and help stakeholders understand the relationship between drought resistance and human thermal stress mitigation for different tree species

This research examined the mechanisms by which temperature, water availability and nitrogen (N) affect the dry matter (DM) yield potential of cocksfoot (Dactylis glomerata L.) dominant pastures. The experiment was a split plot design with main plots of fully irrigated (I) or dryland (D), sub-plots of N fertiliser at 800 kg N/ha in 2003/04; and 1600 kg N/ha in 2004/05 (+N) or 0 kg N/ha (-N). The potential environmental yield of an established 8 year old cocksfoot dominant pasture was 21.9 t DM/ha/y from I+N pastures compared with 9.8 t DM/ha by I-N pastures and 15.1 t DM/ha/y by D+N pastures. The lowest yields were from dryland pastures with no N which produced 7.5 t DM/ha/y in 2003/03 and 5.0 t DM/ha/y in 2004/05. The effect of seasonal temperatures on the DM production, when periods of water stress were excluded, was quantified using thermal time accumulated above a base temperature of 3°C as 7.0 kg DM/°Cd/ha for N fertilised pastures and 3.3 kg DM/°Cd/ha for pastures with no N. The 2.5 t DM/ha difference in yields of D-N pastures in 2003/04 and 2004/05 was the result of the duration, extent and timing of the water stress period. In both years the critical limiting deficit (DL) was calculated as 78 mm from the soil moisture deficit in the 0-0.8 m soil layers. Beyond DL yield decreased at a rate of 1.45%/mm in +N and –N pastures, relative to fully irrigated control pastures. Yields of D+N and D-N pastures were similar during periods of water stress with 0.4±0.1 t/DM/ha produced during the rotation ending 30/12/2003. This was less than from either the I-N (1.2 t DM/ha) or I+N (3.5 t DM/ha) pastures due to the reduction in the amount of photosynthetically active radiation intercepted by the canopies of the dryland pastures. However, in the rotation ending 2/5/2004, after autumn rain alleviated drought conditions, yield of the D+N pasture was 2.1 t DM/ha compared with 1.7 t DM/ha by I+N pastures. The effect of N on yield was described using a nutrition index which showed that as DM yield increased N% in the herbage declined. This is a function of the ratio between metabolic and structural N requirements rather than caused by ontogeny alone. Specific leaf N was determined at two harvests and appeared constant at a given point in time (1.0-1.6 g N/m² leaf). In contrast, specific pseudostem N increased from 0.8-1.0 g N/m² pseudostem at an NNI of 0.4 in –N pastures to 2.6-3.0 g N/m² pseudostem at an NNI of 1.2 in the +N pastures. Differences between the yields of +N and –N pastures were caused by differences in radiation use efficiency (RUE) as determined by the linear relationship (R²=0.76) between RUE and the nitrogen nutrition index (NNI). In this thesis, empirical relationships for the effects of temperature, water availability and N were derived and the physiological mechanisms which underlie these descriptions were identified. These relationships provide clear and simple explanations of the effects of environmental variables on the productivity of cocksfoot based pastures which will enhance understanding of the benefits and limitations of cocksfoot, particularly in dryland farming systems.

O governo do Estado de São Paulo, na década de 60, realizou um programa de introdução de florestas do gênero Pinus para consolidar esta cultura no estado. Algumas destas florestas foram implantadas em áreas do Cerrado, que estavam degradadas devido à utilização para pastagens. Com a intenção de contribuir para um melhor entendimento das características hidrometeorológicas em uma floresta de P. elliotii, os objetivos desta pesquisa foram analisar a repartição da precipitação, o balanço de radiação acima do dossel, o perfil da temperatura do ar e comparar algumas características hidrometeorológicas desta floresta com as características em uma área aberta (gramado) e em uma área campestre de cerrado (pastagem). Na floresta foram instaladas estações meteorológicas automáticas no alto de uma torre e no interior. A repartição da precipitação resultou, pela medição manual, em uma precipitação interna de 75,2%, um escoamento pelo tronco de 0,5% e uma interceptação de 24,3% na floresta. Pela medição automática obteve-se uma precipitação interna de 78,7%. O albedo da floresta apresentou um valor médio de 12,8% e a radiação líquida variou de 6,0 MJm-2dia-1 a 14,2 MJm-2dia-1\'. A diferença da temperatura média do ar, no dossel e no interior da floresta, foi de 0,4ºC. Esta pesquisa apresenta uma grande contribuição para o cenário nacional na área de monitoramento hidrometeorológico de florestas.
The government of São Paulo State, Brazil, forty years ago, carried out a program of introduction of forest species with the main objective to consolidate the Pinus culture in the State. Some of these forests were implanted in areas of the Cerrado that were degraded due their use for grassland. The aim of the present work was to verify the hydrometeorologycal characteristics of Pinus elliottii forest, located in Cerrado, in the central region of São Paulo State, and to compare some characteristics among forest, grassland and lawn. In the forest were installed two automatic meteorological stations, one on the tower top and other inside of the forest. The partitionings of the precipitation by manual measurement were 75.2% for throughfall, 0.5% for stemflow and 24.3% for forest interception. By automatic measurement it was obtained 78.7% for throughfall. Forest\'s albedo presented an average value of 12.8% and net radiation ranged from 6.0 MJm-2day-1 to 14.2 MJm-2day-1. The diference of average air temperature between the canopy and the interior of the forest was 0.4ºC. This study presents a great contribuition to the national ambit of the hydrometeorologycal monitoring in forests.

Conselho Nacional de Desenvolvimento CientÃfico e TecnolÃgico
A pesquisa foi realizada com o objetivo de avaliar o crescimento e as caracterÃsticas morfofisiolÃgicas do dossel de Panicum maximum cv. TanzÃnia sob lotaÃÃo rotativa com trÃs perÃodos de descanso e dois resÃduos pÃs-pastejo. A pesquisa foi conduzida no campo avanÃado do NÃcleo de Ensino e Estudo em Forragicultura-NEEF/DZ/CCA/UFC, localizado na Fazenda Experimental Vale do CurÃ-FEVC/CCA/UFC, em Pentecoste/CE. O mÃtodo de pastejo utilizado foi o de lotaÃÃo rotativa com taxa de lotaÃÃo variÃvel com perÃodo de pastejo de quatro dias. O delineamento foi o inteiramente casualizado em um arranjo fatorial 3 x 2 sendo, trÃs perÃodos de descanso do pasto estabelecidos em funÃÃo do tempo necessÃrio para a restauraÃÃo de um Ãndice de Ãrea foliar que intercepte 85; 95 e 97% da radiaÃÃo fotossinteticamente ativa (RFA) e dois resÃduos pÃs-pastejo, constituÃdos de dois IAFâs residuais, 1,0 e 1,8. Tais variÃveis foram determinadas por meio do Analisador PAR/LAI em Agricultura DECAGON PAR-80 (DECAGON Devices, Inc., Pullman, Washington-USA). O intervalo entre pastejo foi maior para PD 97% IRFA (34 dias) e para o IAFr 1,0 (28,2 dias). A curva de evoluÃÃo da IRFA apresentou-se assintÃtica em relaÃÃo a idade de rebrotaÃÃo do capim, estabilizando-se ao nÃvel de 95% IRFA. JÃ a curva de evoluÃÃo do IAF foi crescente, mesmo apÃs atingir o IAF crÃtico. Houve efeito da IRFA, do IAFr (P<0,05) e da interaÃÃo IRFA x IAFr (P<0,15) sobre a TAlF1/TAlF2. Para a TAlF observou-se efeito da IRFA e do IAFr (P<0,05), sendo que a menor interceptaÃÃo (85% IRFA) e o maior IAFr (1,8) foram os que obtiveram maiores TAlF. A TAlH sofreu efeito apenas do PD (P<0,05). Houve efeito da IRFA e do IAFr (P<0,05) sobre TST, sendo que a maior interceptaÃÃo (1,20 cm/perxdia) e o IAFr 1,0 (0,96 cm/perxdia) que tiveram a maior TST. A percentagem de 85% IRFA (0,12 folha/dia), assim como o IAFr 1,8 (0,11 folha/dia) obtiveram os maiores valores de TApF. O Filocrono sofreu efeito da IRFA (P<0,05) e da interaÃÃo IRFA x IAFr (P<0,15) com 11,05 dias/folha para 95% IRFA e a interaÃÃo ocorrendo para 85% IRFA. Para TAF houve efeito apenas da IRFA (P<0,05). Houve efeito da IRFA (P<0,05) sobre o IAF. A Altura sofreu efeito tanto da IRFA quanto do IAFr (P<0,05). A MSFT e a MSFV sofreram efeito apenas da IRFA (P<0,05). A MSFM sofreu efeito apenas da interaÃÃo IRFA x IAFr (P<0,15), ocorrendo na interceptaÃÃo de 85%. Houve efeito da interaÃÃo IRFA x IAFr sobre a relaÃÃo MV/MM (P<0,15). Houve efeito apenas do IRFA (P<0,05) sobre a MSLV e MSCV. Houve efeito do IAFr (P<0,05) sobre a relaÃÃo Folha/colmo. A DPP sofreu efeito da IRFA, dos IAFrs (P<0,05) e da interaÃÃo IRFA x IAFr (P<0,15). Houve efeito apenas da IRFA (P<0,05) sobre o nÃmero de Folhas/perfilho. No que tange as variÃveis estruturais pÃs pastejo observou-se um efeito do IAFr (P<0,05) sobre a IRFA. A Altura residual sofreu apenas efeito do IAFr (P<0,05). A MSFT e a MSFM sofreram efeito da IRFA, dos IAFrs (P<0,05) e da interaÃÃo IRFA x IAFr (P<0,15). A MSFV e a MSLV sofreram efeito dos IAFrs (P<0,05). Houve interaÃÃo entre o IRFA x IAFr (P<0,15) para a relaÃÃo MV/MM. Houve efeito da IRFA e IAFrs (P<0,05) sobre a MSCV. Houve efeito da IRFA (P<0,05) sobre a relaÃÃo Folha/colmo. A DPP apresentou efeito da IRFA, dos IAFrs (P<0,05) e da interaÃÃo IRFA x IAFr (P<0,15). O nÃmero de Folhas/perfilho estimado pelo mÃtodo morfogÃnico subestimou os valores obtidos pelo mÃtodo visual de campo, mas sendo que apresentou valores confiÃveis e na mesma tendÃncia do nÃmero de Folhas/perfilho obtida pelo mÃtodo visual. O mesmo observa-se para os mÃtodos de obtenÃÃo dos valores da DPP onde o mÃtodo do quadrado superestimou o mÃtodo do transceto, mas apresentando valores dentro da realidade da forrageira e do mÃtodo de pastejo utilizado, podendo perfeitamente ser empregado, facilitando a obtenÃÃo dos valores dessa variÃvel. Valores da TPF e da TAF estimados pelo mÃtodo agronÃmico superestimaram os valores obtidos pelo mÃtodo morfogÃnico.
The research was carried through with the objective of evalua ting the growth and morphophysiological characteristics in canopy of Tanzania grasses under intermittent grazing with three defoliation frequencies and two post-grazing residues. The research was driven in the advanced field of the Nucleu s of Teaching and Study in Forragicultura- NEEF/DZ/CCA/UFC, located in Experi mental Farme it is Worth of CurÃ- FEVC/CCA/UFC, in Pentecoste/CE. The method of used grazing was it of intermittent grazing with tax of variable capacity with peri od of pastejo of four days. The split plot was it entirely randomizer in a factorial arrangemen t 3 x 2 being, three defoliation frequencies of the pasture established in f unction of the necessary time fo r the restoration of an area index to foliate that it intercepts 85; 95 a nd 97% of the photosyntheti cally active radiation (PAR) and two residues powder grazing, constitu ted of two residual IAF's, 1.0 and 1.8. Such variables were certain through the An alisador PAR/LAI in Agricultura DECAGON PAR-80 (DECAGON Devices, Inc ., Pullman, Washington-use s). The interval among pastejo went larger for PD 97% IRFA (34 days) and for IAFr 1.0 (28.2 days). The curve of evolution of IRFA came asymptot ic in relation to age of re growth of the grass, being stabilized at the level of 95% interception PA R. Already the curve of evolution of IAF was growing, even after reaching critical IAF. Ther e was effect of IRFA, of IAFr (P<0.05) and of the interaction IRFA x IAFr (P<0.15) on TAlF1/TAlF2. Para TAlF was noticed effect of IRFA and of IAFr (P<0.05), a nd smallest interception (85% IRFA) and largest IAFr (1.8) they were the ones that obtaine d larger TAlF. TAlH suffered e ffect just of IRFA (P<0.05). There was effect of IRFA and of IAFr (P<0.05) on TST, and most lingering IRFA (1.20 cm/perxdia) and IAFr 1.0 (0.96 cm/perxdia) that had largest TST. The interception of 85% (0.12 leaf/days), as well as IAFr 1.8 (0.11 l eaf/days) they obtaine d the largest values of TApF. Phylochron suffered e ffect of IRFA (P<0.05) and of the interaction IRFA x IAFr (P<0.15) with 11.05 days/leaf for 95% IRFA an d the interaction happening for 85% IRFA. There was effect just of IRFA (P<0.05) for TAF. There was e ffect of IRFA (P <0.05) on IAF. The height suffered so much effect of IRFA, as of IAFr (P<0.05). MSFT and MSFV suffered effect just of IRFA (P<0.05). MSFM suff ered effect just of the interac tion IRFA x IAFr (P<0.15), happening for 85% IRFA. There was effect of the interaction IRFA x IAFr about the relationship MV/MM (P<0.15). There was effect just of IRFA (P<0.05) on MSLV and MSCV. There was effect of IAFr (P<0.05) about the relationship Leaf/stem.

CUTRIM JÚNIOR, José Antônio Alves. Crescimento e morfofisiologia do dossel do capim-tanzânia com três freqüências de desfolhação e dois resíduos pós-pastejo. 2007. 104 f. : Dissertação (mestrado) - Universidade Federal do Ceará, Centro de Ciências Agrárias, Departamento de Zootecnia, Fortaleza-CE, 2007
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The research was carried through with the objective of evalua ting the growth and morphophysiological characteristics in canopy of Tanzania grasses under intermittent grazing with three defoliation frequencies and two post-grazing residues. The research was driven in the advanced field of the Nucleu s of Teaching and Study in Forragicultura- NEEF/DZ/CCA/UFC, located in Experi mental Farme it is Worth of Curú- FEVC/CCA/UFC, in Pentecoste/CE. The method of used grazing was it of intermittent grazing with tax of variable capacity with peri od of pastejo of four days. The split plot was it entirely randomizer in a factorial arrangemen t 3 x 2 being, three defoliation frequencies of the pasture established in f unction of the necessary time fo r the restoration of an area index to foliate that it intercepts 85; 95 a nd 97% of the photosyntheti cally active radiation (PAR) and two residues powder grazing, constitu ted of two residual IAF's, 1.0 and 1.8. Such variables were certain through the An alisador PAR/LAI in Agricultura DECAGON PAR-80 (DECAGON Devices, Inc ., Pullman, Washington-use s). The interval among pastejo went larger for PD 97% IRFA (34 days) and for IAFr 1.0 (28.2 days). The curve of evolution of IRFA came asymptot ic in relation to age of re growth of the grass, being stabilized at the level of 95% interception PA R. Already the curve of evolution of IAF was growing, even after reaching critical IAF. Ther e was effect of IRFA, of IAFr (P<0.05) and of the interaction IRFA x IAFr (P<0.15) on TAlF1/TAlF2. Para TAlF was noticed effect of IRFA and of IAFr (P<0.05), a nd smallest interception (85% IRFA) and largest IAFr (1.8) they were the ones that obtaine d larger TAlF. TAlH suffered e ffect just of IRFA (P<0.05). There was effect of IRFA and of IAFr (P<0.05) on TST, and most lingering IRFA (1.20 cm/perxdia) and IAFr 1.0 (0.96 cm/perxdia) that had largest TST. The interception of 85% (0.12 leaf/days), as well as IAFr 1.8 (0.11 l eaf/days) they obtaine d the largest values of TApF. Phylochron suffered e ffect of IRFA (P<0.05) and of the interaction IRFA x IAFr (P<0.15) with 11.05 days/leaf for 95% IRFA an d the interaction happening for 85% IRFA. There was effect just of IRFA (P<0.05) for TAF. There was e ffect of IRFA (P <0.05) on IAF. The height suffered so much effect of IRFA, as of IAFr (P<0.05). MSFT and MSFV suffered effect just of IRFA (P<0.05). MSFM suff ered effect just of the interac tion IRFA x IAFr (P<0.15), happening for 85% IRFA. There was effect of the interaction IRFA x IAFr about the relationship MV/MM (P<0.15). There was effect just of IRFA (P<0.05) on MSLV and MSCV. There was effect of IAFr (P<0.05) about the relationship Leaf/stem.
A pesquisa foi realizada com o objetivo de avaliar o crescimento e as características morfofisiológicas do dossel de Panicum maximum cv. Tanzânia sob lotação rotativa com três períodos de descanso e dois resíduos pós-pastejo. A pesquisa foi conduzida no campo avançado do Núcleo de Ensino e Estudo em Forragicultura-NEEF/DZ/CCA/UFC, localizado na Fazenda Experimental Vale do Curú-FEVC/CCA/UFC, em Pentecoste/CE. O método de pastejo utilizado foi o de lotação rotativa com taxa de lotação variável com período de pastejo de quatro dias. O delineamento foi o inteiramente casualizado em um arranjo fatorial 3 x 2 sendo, três períodos de descanso do pasto estabelecidos em função do tempo necessário para a restauração de um índice de área foliar que intercepte 85; 95 e 97% da radiação fotossinteticamente ativa (RFA) e dois resíduos pós-pastejo, constituídos de dois IAF’s residuais, 1,0 e 1,8. Tais variáveis foram determinadas por meio do Analisador PAR/LAI em Agricultura DECAGON PAR-80 (DECAGON Devices, Inc., Pullman, Washington-USA). O intervalo entre pastejo foi maior para PD 97% IRFA (34 dias) e para o IAFr 1,0 (28,2 dias). A curva de evolução da IRFA apresentou-se assintótica em relação a idade de rebrotação do capim, estabilizando-se ao nível de 95% IRFA. Já a curva de evolução do IAF foi crescente, mesmo após atingir o IAF crítico. Houve efeito da IRFA, do IAFr (P<0,05) e da interação IRFA x IAFr (P<0,15) sobre a TAlF1/TAlF2. Para a TAlF observou-se efeito da IRFA e do IAFr (P<0,05), sendo que a menor interceptação (85% IRFA) e o maior IAFr (1,8) foram os que obtiveram maiores TAlF. A TAlH sofreu efeito apenas do PD (P<0,05). Houve efeito da IRFA e do IAFr (P<0,05) sobre TST, sendo que a maior interceptação (1,20 cm/perxdia) e o IAFr 1,0 (0,96 cm/perxdia) que tiveram a maior TST. A percentagem de 85% IRFA (0,12 folha/dia), assim como o IAFr 1,8 (0,11 folha/dia) obtiveram os maiores valores de TApF. O Filocrono sofreu efeito da IRFA (P<0,05) e da interação IRFA x IAFr (P<0,15) com 11,05 dias/folha para 95% IRFA e a interação ocorrendo para 85% IRFA. Para TAF houve efeito apenas da IRFA (P<0,05). Houve efeito da IRFA (P<0,05) sobre o IAF. A Altura sofreu efeito tanto da IRFA quanto do IAFr (P<0,05). A MSFT e a MSFV sofreram efeito apenas da IRFA (P<0,05). A MSFM sofreu efeito apenas da interação IRFA x IAFr (P<0,15), ocorrendo na interceptação de 85%. Houve efeito da interação IRFA x IAFr sobre a relação MV/MM (P<0,15). Houve efeito apenas do IRFA (P<0,05) sobre a MSLV e MSCV. Houve efeito do IAFr (P<0,05) sobre a relação Folha/colmo. A DPP sofreu efeito da IRFA, dos IAFrs (P<0,05) e da interação IRFA x IAFr (P<0,15). Houve efeito apenas da IRFA (P<0,05) sobre o número de Folhas/perfilho. No que tange as variáveis estruturais pós pastejo observou-se um efeito do IAFr (P<0,05) sobre a IRFA. A Altura residual sofreu apenas efeito do IAFr (P<0,05). A MSFT e a MSFM sofreram efeito da IRFA, dos IAFrs (P<0,05) e da interação IRFA x IAFr (P<0,15). A MSFV e a MSLV sofreram efeito dos IAFrs (P<0,05). Houve interação entre o IRFA x IAFr (P<0,15) para a relação MV/MM. Houve efeito da IRFA e IAFrs (P<0,05) sobre a MSCV. Houve efeito da IRFA (P<0,05) sobre a relação Folha/colmo. A DPP apresentou efeito da IRFA, dos IAFrs (P<0,05) e da interação IRFA x IAFr (P<0,15). O número de Folhas/perfilho estimado pelo método morfogênico subestimou os valores obtidos pelo método visual de campo, mas sendo que apresentou valores confiáveis e na mesma tendência do número de Folhas/perfilho obtida pelo método visual. O mesmo observa-se para os métodos de obtenção dos valores da DPP onde o método do quadrado superestimou o método do transceto, mas apresentando valores dentro da realidade da forrageira e do método de pastejo utilizado, podendo perfeitamente ser empregado, facilitando a obtenção dos valores dessa variável. Valores da TPF e da TAF estimados pelo método agronômico superestimaram os valores obtidos pelo método morfogênico.

FundaÃÃo Cearense de Apoio ao Desenvolvimento Cientifico e TecnolÃgico
A pesquisa foi conduzida no campo avanÃado do NÃcleo de Ensinos e Estudos em Forragicultura-NEEF/DZ/CCA/UFC (www.neef.ufc.br), localizado na fazenda experimental Vale do Curu â FEVC/CCA/UFC, em Pentecoste/CE. Objetivou-se avaliar a composiÃÃo quÃmico-bromatolÃgica e digestibilidade da dieta consumida e o desempenho produtivo de ovinos pastejando em Panicum maximum cv. tanzÃnia sob trÃs freqÃÃncias de desfolhaÃÃo determinadas pela interceptaÃÃo da radiaÃÃo fotossintÃticamente ativa (IRFA) de 85, 95 e 97% e dois resÃduos pÃs-pastejo, determinados pelos Ãndices de Ãrea foliar (IAF) residuais de 1,0 e 1,8. O delineamento experimental foi inteiramente casualizado com arranjo fatorial 3x2 (trÃs freqÃÃncias de desfolhaÃÃo e dois resÃduos pÃs pastejo), com quatro repetiÃÃes para as variÃveis do pasto (piquetes) e seis para as variÃveis relacionadas aos animais (ovinos). Foram utilizados 90 ovinos sem padrÃo racial definido (SPRD), com peso vivo mÃdio inicial de 20,0 kg. Em cada tratamento foram utilizados 6 ovinos, resultando em 36 animais de prova. Os demais ovinos foram utilizados como animais de equilÃbrio, quando necessÃrio para rebaixar o pasto para o IAF residual preconizado para aquele tratamento, ao final do 4Â dia de pastejo. Para determinaÃÃo da composiÃÃo quÃmico-bromatolÃgica, digestibilidade e ensaio de cinÃtica de degradaÃÃo da matÃria seca da dieta, foi realizada uma simulaÃÃo de pastejo no 1Â e 4Â dia de ocupaÃÃo. Foi avaliado ainda o ensaio de desempenho animal, alÃm da determinaÃÃo da capacidade de suporte em cada sistema sob lotaÃÃo rotativa. A diminuiÃÃo da freqÃÃncia de desfolhaÃÃo reduziu o valor nutritivo do capim-tanzÃnia e o desempenho dos ovinos. Maior rendimento de peso vivo animal por Ãrea ao longo do ano foi obtido na freqÃÃncia de desfolhaÃÃo de 85% nos dois resÃduos pÃs-pastejo e na freqÃÃncia de desfolhaÃÃo intermediÃria, apenas no Ãndice de Ãrea foliar residual de 1,0. Portanto, recomenda-se adotar uma freqÃÃncia de esfolhaÃÃo no ix capim-tanzÃnia menor ou igual a 95% de interceptaÃÃo da radiaÃÃo fotossinteticamente ativa, com um IAF residual de 1,0, a fim de compatibilizar os objetivos de produÃÃo animal e manutenÃÃo da qualidade do pasto no longo prazo.
The research was carried out at the advanced field of the Forage Teaching and Research Nucleus - NEEF/DZ/CCA/UFC (www.neef.ufc.br), located at Curu Valley Experimental Farm - FEVC/CCA/UFC, in Pentecoste/CE. The aim of this research was to evaluate the nutritional value of the simulated diet and the productive performance of sheep grazing in Panicum maximum cv. Tanzania under three defoliation frequencies, determined by the interception of the photosynthetically active radiation in canopy (PAR) of 85, 95 and 97 % and two post-grazing residues, determined by the residual leaf area index (LAI) of 1.0 and 1.8. A completely randomized design in factorial arrangement 3x2 (three defoliation frequencies and two residues after graze), with four repetitions for the variable of the grass (poles) and around the six (sheep) for the variable related to the animals, having still cycles of graze as measured repeated in the time. 90 sheep without defined breeding standard had been used (SPRD), averaging an initial live weight of 20.0 kg were used. 6 sheep were allocated to each treatment, resulting in 36 tester sheep. The others 44 sheep were used as put-and-taken animal, to keep the canopy as established by the end of the grazing period. For determination of the chemical composition assay, digestibility and an in situ dry matter degradation trial, a simulation of grazing in 1st and 4th days was carried out. An animal performance assay was conducted, beyond the determination of the carrying capacity in each grazing system. The defoliation frequency reduction diminished the diet nutritional value and the sheep performance. The highest animal live weight productivity toward the year was obtained in the 85% defoliation frequency on both post-grazing residues and at the intermediate defoliation frequency only at the residual leaf area index of 1.0. So, one xi recommends to adopt a defoliation frequency of tanzania grass up to 95% of photosynthetically active radiation interception, with a residual LAI of 1.0 to obtain a compromise between the objectives of animal production and forage quality maintenance in the long term.

Conselho Nacional de Desenvolvimento CientÃfico e TecnolÃgico
Avaliou-se os componentes da biomassa e a estrutura do dossel em capim-aruana sob duas frequÃncias (FD - nÃveis de interceptaÃÃo de 85 e 95% da radiaÃÃo fotossinteticamente ativa incidente no topo do dossel) e duas intensidades (ID - Ãndices de Ãrea foliar residual de 1,0 e 1,8) de desfolhaÃÃo, em um delineamento inteiramente casualizado, num arranjo fatorial 2x2, com cinco repetiÃÃes (piquetes). O experimento foi conduzido no campo avanÃado do NÃcleo de Ensino e Estudos em Forragicultura, situado na Fazenda experimental Vale do CurÃ, localizada no municÃpio de Pentecoste-CE. Os momentos de entrada e saÃda dos animais foram determinados utilizando-se o Analisador PAR/LAI em Agricultura DECAGON PAR-80 para as leituras da interceptaÃÃo de radiaÃÃo fotossinteticamente ativa (IRFA) e do Ãndice de Ãrea foliar (IAF). Foram avaliadas no inÃcio e no final do perÃodo de descanso de cada um dos cinco piquetes experimentais, as seguintes caracterÃsticas estruturais: altura do pasto; altura da primeira lÃgula viva, a mais velha abaixo do dossel; horizonte de pastejo e densidade populacional de perfilhos. Os componentes da biomassa de forragem total (BFT) foram quantificados no inÃcio e final do perÃodo de descanso, a partir da colheita com duas molduras de 0,25 mÂ, sendo encaminhadas ao laboratÃrio para separaÃÃo dos componentes: folha, colmo e material morto. ApÃs a separaÃÃo dos componentes da BFT, as amostras foram levadas à estufa de ventilaÃÃo forÃada à 55 ÂC atà peso constante para estimativa das seguintes variÃveis: biomassa de forragem total (BFT kg∙ha-1∙ciclo-1), de forragem morta (BFM kg∙ha-1∙ciclo-1 ), de forragem verde (BFV kg∙ha-1∙ciclo-1), de lÃmina foliar verde (BLV kg∙ha-1∙ciclo-1), de colmo verde (BCV kg∙ha-1∙ciclo-1), alÃm das relaÃÃes material vivo/material morto (MV/MM) e folha/colmo (F/C). O maior tempo de descanso foi verificado para os pastos manejados com menor IAFr e menor frequÃncia de desfolhaÃÃo (95% de IRFA), com mÃdia de 24 dias. Os componentes da biomassa e as caracterÃsticas estruturais na condiÃÃo prÃ-pastejo foram afetados (P<0,05) pelas FDs, onde a menor frequÃncia de desfolhaÃÃo proporcionou maiores produÃÃes, com considerÃvel contribuiÃÃo de biomassa de colmos verdes e de forragem morta. A altura prÃ-pastejo apresentou boa correlaÃÃo com a %IRFA na condiÃÃo prÃ-pastejo (r=0,9285; P=0,0001). Pastos manejados com menor FD proporcionaram baixa F/C, com mÃdia de 1,65, enquanto pastos manejados com maior FD apresentaram mÃdia de 2,40. Com o aumento da %IRFA, foram verificados maiores horizontes de pastejo, com mÃdias de 20,2 e 14,2 cm para as frequÃncias de 95 e 85%IRFA, respectivamente. Os componentes da biomassa residual foram afetados (P<0,05) pelas IDs, com maior biomassa de forragem residual observada para os pastos manejados com IAFr de 1,8. Os pastos manejados com 95x1,8 apresentaram BCVr e BFMr superiores (P<0,05) aos demais manejos. Pastos manejados com 95x1,0 apresentaram incremento (P<0,05) de BFT ao longo de ciclos sucessivos de pastejo. O capim-aruana, para utilizaÃÃo com ovinos, deve ser manejado com frequÃncia de 85% de IRFA e intensidade de desfolhaÃÃo equivalente a IAFr de 1,0.
This study evaluated the components of biomass and the canopy structure of aruana grass managed under two frequencies (FD, interception of the photosynthetically active radiation at 85 and 95% incident on the top of the canopy) and two intensities (ID, residual leaf area index of 1.0 and 1.8) of defoliation in a completely randomized design in a 2x2 factorial arrangement with five replications (paddocks). The experiment was conducted at the advanced field of Teaching and Study Center Forage, located in the municipality of Pentecoste, Cearà State. The times of entry and exit of animals were determined using the PAR/LAI Analyzer in Agriculture DECAGON PAR-80 for reading photosynthetically active radiation interception (PARI) and leaf area index (LAI). At the beginning and end of the rest period of each of the five experimental paddocks, the following structural characteristics were evaluated: sward height (cm); height of the first ligule, the oldest below the canopy; grazing horizon and tiller population density. The components of total forage biomass (TFB ha-1 cycle-1) were also quantified at the beginning and end of the rest period, from the collection using two frames of 0.25 mÂ, and sent to the laboratory to separate the components: leaf, stem and dead material. After separation of the TFB components, samples were dried to constant weight in a forced ventilation oven at 55ÂC to constant weight to estimate the following parameters: total forage biomass (TFB kg ha-1 cycle-1), dead forage biomass (DFB kg ha-1 cycle-1), green forage biomass (GFB kg kg ha-1 cycle-1), green leaf biomass (GLB kg ha-1 cycle-1), green stem biomass (GSB kg ha-1 cycle-1), and green/dead material ratio (GM/DM) and leaf/stem ratio (L/S). The longest rest period was observed in pastures managed with lower rLAI and lower defoliation frequency (95% PARI), with average of 24 days. Biomass components and structural characteristics in the pre-grazing condition were affected (P <0.05) by FD, where the lower defoliation frequency resulted in higher production with considerable contribution of biomass of green stems and dead forage. The pre-grazing height showed a high correlation with the %PARI in the pre-grazing condition (r = 0.9285, P = 0.0001). Pastures managed with lower FD had low L/S, with average of 1.65, while those managed with higher FD, an average of 2.40. With increasing %PARI, we verified higher grazing horizons, with averages of 20.2 and 14.2 cm, for 95 and 85 %PARI, respectively. The components of the residual biomass were affected (P <0.05) by ID, with higher residual forage biomass observed for pastures managed with rLAI of 1.8. Pastures managed with 95x1.8 exhibited higher rGSB Âand rDFB (P <0.05) compared to the other managements. Pastures managed with 95x1.0 showed an increase in TFB (P <0.05) over successive grazing cycles. Aruana grass supplied to sheep should be managed under 85% PARI frequency and defoliation intensity equal to 1.0 rLAI.

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SILVA, Leane Veras da. Componentes da biomassa e estrutura do capim-aruana sob duas frequências e duas intensidades de desfolhação. 2015. 56 f . : Dissertação (mestrado) - Universidade Federal do Ceará, Centro de Ciências Agrárias, Departamento de Zootecnia, Fortaleza-CE, 2015
This study evaluated the components of biomass and the canopy structure of aruana grass managed under two frequencies (FD, interception of the photosynthetically active radiation at 85 and 95% incident on the top of the canopy) and two intensities (ID, residual leaf area index of 1.0 and 1.8) of defoliation in a completely randomized design in a 2x2 factorial arrangement with five replications (paddocks). The experiment was conducted at the advanced field of Teaching and Study Center Forage, located in the municipality of Pentecoste, Ceará State. The times of entry and exit of animals were determined using the PAR/LAI Analyzer in Agriculture DECAGON PAR-80 for reading photosynthetically active radiation interception (PARI) and leaf area index (LAI). At the beginning and end of the rest period of each of the five experimental paddocks, the following structural characteristics were evaluated: sward height (cm); height of the first ligule, the oldest below the canopy; grazing horizon and tiller population density. The components of total forage biomass (TFB ha-1 cycle-1) were also quantified at the beginning and end of the rest period, from the collection using two frames of 0.25 m², and sent to the laboratory to separate the components: leaf, stem and dead material. After separation of the TFB components, samples were dried to constant weight in a forced ventilation oven at 55°C to constant weight to estimate the following parameters: total forage biomass (TFB kg ha-1 cycle-1), dead forage biomass (DFB kg ha-1 cycle-1), green forage biomass (GFB kg kg ha-1 cycle-1), green leaf biomass (GLB kg ha-1 cycle-1), green stem biomass (GSB kg ha-1 cycle-1), and green/dead material ratio (GM/DM) and leaf/stem ratio (L/S). The longest rest period was observed in pastures managed with lower rLAI and lower defoliation frequency (95% PARI), with average of 24 days. Biomass components and structural characteristics in the pre-grazing condition were affected (P <0.05) by FD, where the lower defoliation frequency resulted in higher production with considerable contribution of biomass of green stems and dead forage. The pre-grazing height showed a high correlation with the %PARI in the pre-grazing condition (r = 0.9285, P = 0.0001). Pastures managed with lower FD had low L/S, with average of 1.65, while those managed with higher FD, an average of 2.40. With increasing %PARI, we verified higher grazing horizons, with averages of 20.2 and 14.2 cm, for 95 and 85 %PARI, respectively. The components of the residual biomass were affected (P <0.05) by ID, with higher residual forage biomass observed for pastures managed with rLAI of 1.8. Pastures managed with 95x1.8 exhibited higher rGSB ¬and rDFB (P <0.05) compared to the other managements. Pastures managed with 95x1.0 showed an increase in TFB (P <0.05) over successive grazing cycles. Aruana grass supplied to sheep should be managed under 85% PARI frequency and defoliation intensity equal to 1.0 rLAI.
Avaliou-se os componentes da biomassa e a estrutura do dossel em capim-aruana sob duas frequências (FD - níveis de interceptação de 85 e 95% da radiação fotossinteticamente ativa incidente no topo do dossel) e duas intensidades (ID - índices de área foliar residual de 1,0 e 1,8) de desfolhação, em um delineamento inteiramente casualizado, num arranjo fatorial 2x2, com cinco repetições (piquetes). O experimento foi conduzido no campo avançado do Núcleo de Ensino e Estudos em Forragicultura, situado na Fazenda experimental Vale do Curú, localizada no município de Pentecoste-CE. Os momentos de entrada e saída dos animais foram determinados utilizando-se o Analisador PAR/LAI em Agricultura DECAGON PAR-80 para as leituras da interceptação de radiação fotossinteticamente ativa (IRFA) e do índice de área foliar (IAF). Foram avaliadas no início e no final do período de descanso de cada um dos cinco piquetes experimentais, as seguintes características estruturais: altura do pasto; altura da primeira lígula viva, a mais velha abaixo do dossel; horizonte de pastejo e densidade populacional de perfilhos. Os componentes da biomassa de forragem total (BFT) foram quantificados no início e final do período de descanso, a partir da colheita com duas molduras de 0,25 m², sendo encaminhadas ao laboratório para separação dos componentes: folha, colmo e material morto. Após a separação dos componentes da BFT, as amostras foram levadas à estufa de ventilação forçada à 55 °C até peso constante para estimativa das seguintes variáveis: biomassa de forragem total (BFT kg∙ha-1∙ciclo-1), de forragem morta (BFM kg∙ha-1∙ciclo-1 ), de forragem verde (BFV kg∙ha-1∙ciclo-1), de lâmina foliar verde (BLV kg∙ha-1∙ciclo-1), de colmo verde (BCV kg∙ha-1∙ciclo-1), além das relações material vivo/material morto (MV/MM) e folha/colmo (F/C). O maior tempo de descanso foi verificado para os pastos manejados com menor IAFr e menor frequência de desfolhação (95% de IRFA), com média de 24 dias. Os componentes da biomassa e as características estruturais na condição pré-pastejo foram afetados (P<0,05) pelas FDs, onde a menor frequência de desfolhação proporcionou maiores produções, com considerável contribuição de biomassa de colmos verdes e de forragem morta. A altura pré-pastejo apresentou boa correlação com a %IRFA na condição pré-pastejo (r=0,9285; P=0,0001). Pastos manejados com menor FD proporcionaram baixa F/C, com média de 1,65, enquanto pastos manejados com maior FD apresentaram média de 2,40. Com o aumento da %IRFA, foram verificados maiores horizontes de pastejo, com médias de 20,2 e 14,2 cm para as frequências de 95 e 85%IRFA, respectivamente. Os componentes da biomassa residual foram afetados (P<0,05) pelas IDs, com maior biomassa de forragem residual observada para os pastos manejados com IAFr de 1,8. Os pastos manejados com 95x1,8 apresentaram BCVr e BFMr superiores (P<0,05) aos demais manejos. Pastos manejados com 95x1,0 apresentaram incremento (P<0,05) de BFT ao longo de ciclos sucessivos de pastejo. O capim-aruana, para utilização com ovinos, deve ser manejado com frequência de 85% de IRFA e intensidade de desfolhação equivalente a IAFr de 1,0.

CoordenaÃÃo de AperfeÃoamento de Pessoal de NÃvel Superior
Avaliou-se as trocas gasosas, o fluxo de biomassa e as caracterÃsticas estruturais do capim-aruana pastejado sob lotaÃÃo rotativa por ovinos Morada Nova submetidos a quatro nÃveis de suplementaÃÃo concentrada (0,0; 0,6; 1,2 e 1,8% do PC), com perÃodo de ocupaÃÃo de quatro dias e descanso quando da interceptaÃÃo de 90% da radiaÃÃo fotossinteticamente ativa (IRFA), num delineamento inteiramente casualizado, num arranjo em parcelas subdivididas, sendo os nÃveis de suplementaÃÃo as parcelas e os ciclos de pastejo as subparcelas. A condiÃÃo de pÃs-pastejo consistiu em um Ãndice de Ãrea foliar de 1,0. Constatou-se efeito linear crescente para a taxa de alongamento das hastes e taxa de aparecimento foliar em funÃÃo dos nÃveis de suplementaÃÃo. As taxas de produÃÃo e acÃmulo de forragem foram incrementadas pelos nÃveis de suplementaÃÃo. As variÃveis das trocas gasosas das folhas recÃm-expandidas: transpiraÃÃo, condutÃncia estomÃtica, taxa de fotossÃntese foliar, mostraram efeito somente de ciclo de pastejo. NÃo houve nenhum efeito significativo sobre as variÃveis das trocas gasosas das folhas emergentes. Os componentes da biomassa pÃs-pastejo registraram efeito significativo da interaÃÃo nÃveis de suplementaÃÃo e ciclos de pastejo para biomassa de forragem verde residual (BFVr), lÃmina foliar verde residual (BLVr), colmo verde residual (BCVr) e relaÃÃo folha:colmo (F_Cr) residual, estas variÃveis, com exceÃÃo da F_Cr apresentaram efeito quadrÃtico nos ciclos dois e trÃs, confirmando o efeito substitutivo, a partir do nÃvel de suplementaÃÃo de 0,86%, com o mÃnimo estimado de 257,53 Kgâha-Ââano-1, para a BLVr. A relaÃÃo material vivo_ material morto residual apresentou efeito linear crescente em funÃÃo dos nÃveis de suplementaÃÃo. A altura do pasto e a densidade populacional de perfilhos residual foram afetadas somente pelos ciclos de pastejo. A biomassa de forragem verde (BFV) e de lÃmina foliar verde (BLV) apresentaram efeito da interaÃÃo nÃveis de suplementaÃÃo e ciclos de pastejo. A BLV registrou efeito quadrÃtico para os ciclos dois e trÃs, onde os maiores valores foram de 1489,47, no nÃvel de 0,62, do segundo ciclo e de 1554,95 Kgâha-Ââano-1no ciclo trÃs com nÃvel de suplementaÃÃo de 0,14 %. As trocas gasosas nÃo sÃo afetadas pelos nÃveis de suplementaÃÃo. As caracterÃsticas morfogÃnicas do pasto de capim-aruana sÃo afetadas linearmente pelos nÃveis crescentes de suplementaÃÃo. Os componentes estruturais do pasto de capim-aruana sÃo afetados pelos nÃveis crescentes de suplementaÃÃo concentrada, registrando efeito substitutivo do suplemento.
This study evaluated the gas exchange, the biomass flow and the structural characteristics of Panicum maximum cv. Aruana grazed by Morada Nova sheep under rotational stocking and supplied with four concentrate supplementation levels (0.0; 0.6; 1.2 and 1.8% body weight), using four day-grazing periods and rest when the interception of 90% of the photosynthetic active radiation (PAR). The experiment was a completely randomized split-plot design with supplementation levels as plots and grazing cycles as subplots. The post-grazing condition consisted of a leaf area index of 1.0. We observed an increasing linear effect for stem elongation rate and leaf appearance rate according to the supplementation levels. The production and accumulation rates of forage also increased according to the supplementation levels. The different grazing cycles affected the variables of gas exchange of the recently expanded leaves, namely transpiration, stomatal conductance, leaf photosynthesis rate. In turn, no significant effect was detected for the gas exchange variables of emergent leaves. Biomass components during the post-grazing period showed a significant effect of the interaction between supplementation levels and grazing cycles for residual green forage biomass (BFVr), residual green leaf biomass (BLVr), residual green stem biomass (BCVr) and residual leaf:stem ratio (F_Cr). All these variables, with the exception of F_Cr, presented a quadratic effect in the cycles two and three, confirming the substitutive effect from the supplementation level of 0.86%, with the minimum estimated at 257.53 Kgâha-Ââyear-1, for BLVr. The ratio between living and senescent material showed an increasing linear response as a function of the supplementation levels. The pasture height and the residual tiller population density were affected only by grazing cycles. Green forage biomass (BFV) and green leaf biomass (BLV) presented a quadratic effect of the supplementation levels and grazing cycles. BLV showed a quadratic effect for the cycles two and three, where the higher values were 1489.47, at the level of 0.62, in the second cycle, and 1554.95 Kgâha-Ââyear-1 in the third cycle with a supplementation level of 0.14%. Gas exchange was not influenced by the supplementation levels. Morphogenetic traits of the aruana grass pasture are linearly affected by increasing levels of supplementation. The structural components of the aruana grass pasture are influenced by the increasing levels of concentrate supplementation, with a substitutive effect of the supplement.

Genotypic differences in growth and yield of two common bean (Phaseolus vulgaris L) cultivars to Rhizobium inoculation and management were investigated. In 2003-04, the two bean cultivars (Scylla and T-49) were combined with three inoculant treatments (strains CC 511 and RCR 3644, and a control of no inoculation), two fertiliser levels (0 and 150 kg N ha⁻¹) and two irrigation treatments (irrigated and rainfed). There was no nodulation on either cultivar. To further investigate the symbiotic relationship, 16 rhizobial isolates, including the two used in the first field experiment, were combined with the cultivar Scylla and evaluated in a greenhouse. Subsequently, five Rhizobium isolates were chosen for further field evaluation, based on signs of early nodulation in the greenhouse trial. The second field experiment in 2004-05 combined the five inoculant strains (RCR 3644, UK 2, H 20, PRF 81, PhP 17 and a control) with two bean cultivars (Scylla and T-49). In the greenhouse, nodule number varied from 7 (UK 2) to 347 (H 441) nodules plant⁻¹ at 51 DAS and from 13 (UK 1) to 335 (CIAT 899) nodules plant⁻¹ at 85 DAS. In 2004-05, in the field, nodulation was also variable, ranging between 1 and approximately 70 nodules plant⁻¹, with higher nodules numbers plant⁻¹ being found on cultivar T-49. Of the isolates used in the field, strains H 20, PRF 81 and PhP 17 produced 70, 25 and 12 nodules plant⁻¹ at 70, 40 and 54 DAS respectively. Nodules formed were of various sizes and more than 80 % were pink to dark red in colour denoting the presence of leghaemoglobin and active N fixation. The remaining nodules were either green or white. The importance of selecting an appropriate cultivar for the growing conditions was highlighted in these experiments. Leaf area index, leaf area duration intercepted radiation and final utilisation efficiency were significantly affected by cultivar. In both seasons cv. T-49 reached maturity (dry seed) before Scylla, while unirrigated plants reached green pod maturity seven days before irrigated plants. Plants of cv. Scylla gave a final TDM of 730 g m⁻²; compared to the 530 g m⁻² produced by T-49. The average growth rate was 7.0 and 5.2 g m⁻² day⁻¹ for Scylla and T-49 respectively (2003-04). Plants receiving 150 kg N ha⁻¹ produced 665 g m⁻² TDM which was 12 % more than was produced by unfertilised plants. The application of 150 kg N ha⁻¹ gave an average growth rate of 6.4 g m⁻² day⁻¹ compared to 5.7 g m⁻² day⁻¹ from plants with no N. Inoculation in the field had no significant effect on TDM in both seasons. Temperature affected growth and DM accumulation. Accumulated DM was highly dependent on cumulative intercepted PAR. Air temperatures below the base temperature (10 °C) affected growth in 2004-05, resulting in plants accumulating just 0.24 g DM MJ⁻¹ PAR during early growth. This increased to 2.26 g DM MJ⁻¹ PAR when the temperature was increased above the base temperature. There was a strong relationship between LAI and intercepted PAR. A LAI of 4.0-4.5 was required to intercept 90-95 % of incident solar radiation. Cultivar significantly (p < 0.001) affected radiation use efficiency (RUE). Scylla had a RUE of 1.02 g DM MJ⁻¹ PAR compared to T-49 at 1.18 g DM MJ⁻¹ PAR. Seed yield was significantly (p < 0.001) affected by cultivar and fertiliser application. Cultivar Scylla produced 467 g m⁻² which was 76 % more than T-49, while a 12 % increase in seed yield was observed in N fertilised plants over unfertilised plants. Only cultivar significantly affected HI, while the yield components that had the greatest effect on seed yield were hundred seed weight and pods plant⁻¹. Inoculation significantly (p< 0.05) affected 100 seed weight (2004-05). Plants inoculated with strain H 20 had the highest 100 seed weight at 25.2 g with cv. Scylla producing larger seeds than T-49. The belief that local environmental conditions play a major role on field survival of bacteria, led to the use of PCR methods to identify field nodulating organisms. Amplification of genomic DNA from parent isolates using primers fC and rD generated a single band for each isolate. Isolates were identified to the species level as either Rhizobium or Agrobacterium, using the highly conserved internally transcribed spacer (ITS) region and are known to nodulate common bean. The DNA extracted from the isolates recovered from nodules of field grown beans gave multiple bands with primers fC and rD. Five distinct banding patterns were observed. All of these were different from those of parent isolates. Sequencing of the 16S rRNA demonstrated that nodules of field grown beans in Canterbury were inhabited by Pseudomonads either alone or in association with other root nodulating organisms. The inability to identify the inoculant strains in nodules of field grown beans does not rule out their infection and nodulating function in the cultivars used. The results suggest the possibility of both Rhizobium and Pseudomonads cohabiting in the nodules of field grown beans. The aggressive nature of Pseudomonads on artificial media, possibly out competing the inoculant rhizobia is proposed, leading to the inability to identify the inoculant strain from the nodules of the field grown beans by PCR methods. The need to identify the nodule forming or nodule inhabiting bacteria in the nodules is necessary to classify the importance of these organisms and their economic benefit to agricultural production. This study also underlines the importance of using PCR methods to gain valuable insights into the ecological behaviour of Rhizobium inoculants and nodule inhabiting organisms.

Cette étude propose d’analyser l’influence de l’architecture du palmier à huile sur sa capacité à intercepter la lumière, en se basant sur des reconstructions 3D de palmiers et en établissant un bilan radiatif sur ses structures végétales reconstruites in silico. Le premier objectif de l’étude était de caractériser et modéliser la variabilité génétique de l’architecture du palmier à huile et de son interception lumineuse. Dans un deuxième objectif l’amélioration potentielle de l’interception de la lumière et de l’assimilation carbonée a été évaluée en modifiant les traits morphologiques et géométriques des feuilles et des idéotypes architecturaux de palmiers à huile ont été proposés.Des relations allométriques ont été utilisées pour modéliser les traits architecturaux en fonction de gradients ontogénétique et de topologie des feuilles dans la couronne. La méthode permet de reconstruire des palmiers à huile virtuels à différents âges au cours du développement. De plus, l’approche allométrique a été couplée à des modèles à effets mixtes pour intégrer au travers de paramètres la variabilité entre et au sein des cinq progénies. Le modèle permet ainsi de simuler les spécificités architecturales des cinq progenies en incluant les variabilités entre individus observés. Le modèle architectural, paramétré pour les différentes progénies, a ensuite été implémenté dans AMAPstudio pour générer des maquettes 3D de palmiers et ainsi estimer leur interception lumineuse, de l’individu à la parcelle entière.Les résultats de ces analyses ont révélé des différences significatives entre et au sein des progenies, dans la géométrie des feuilles (longueur du pétiole, densité de folioles sur le rachis, et courbure du rachis) et dans la morphologie des folioles (gradients de longueurs et largeurs le long du rachis). La comparaison virtuelle des différentes progénies ont aussi montré des efficacités distinctes de l’interception lumineuse.Des analyses de sensibilité ont ensuite été réalisées pour identifier les traits architecturaux influençant l’interception lumineuse et l’assimilation potentielle à différents âges de la plante. Les paramètres les plus sensibles au cours du développement furent ceux reliés à la surface totale foliaire (longueur des rachis, nombre de folioles, morphologie des folioles), mais les attributs géométriques plus fins de la feuille ont montré un effet croissant avec la fermeture de la canopée. Sur un couvert adulte, l’optimum en assimilation carbonée est atteint pour des indices de surfaces foliaires (LAI) entre 3,2 et 5,5 m2.m−2, avec des feuilles érigées, de courts pétioles et rachis et un nombre important de folioles sur le rachis. Quatre idéotypes architecturaux pour l’assimilation carbonée ont été proposés et présentent des combinaisons spécifiques de traits géométriques, limitant l’ombrage mutuel des plantes et optimisant la distribution de la lumière dans la couronne.En conclusion, le modèle 3D de palmiers à huile, dans sa conception et son application, a permis de détecter les traits architecturaux génétiquement déterminés et influençant l’interception lumineuse. Ainsi, le nombre limité de traits dégagés par l’analyse de sensibilité ainsi que les combinaisons de traits révélées au travers des idéotypes pourraient être pris en compte dans de futurs programmes de sélection. En perspective, des travaux dédiés à intégrer dans ce modèle d’autres processus physiologiques, tels que la régulation de la conductance stomatique et le partitionnement du carbone dans la plante, sont à envisager. Ce nouvel FSPM pourrait alors être utilisé pour tester différents scénarii, comme par exemple dans un contexte de changement climatique avec de faibles radiations et des périodes de sécheresse fréquentes. De même, ce modèle pourrait être utilisé pour étudier différentes configurations de plantation et des systèmes de cultures intercalaires, et ainsi proposer de nouveaux idéotypes multicritères
In this study we proposed to investigate the influence of oil palm architecture on the capacity of the plant to intercept light, by using 3D reconstructions and model-assisted evaluation of radiation-use efficiency. The first objective of this study was to analyse and model oil palm architecture and light interception taking into account genetic variability. A second objective was to explore the potential improvements in light capture and carbon assimilation by manipulating oil palm leaf traits and propose architectural ideotypes.Allometric relationships were applied to model these traits according to ontogenetic gradients and leaf position within the crown. The methodology allowed reconstructing virtual oil palms at different stages over plant development. Additionally, the allometric-based approach was coupled to mixed-effect models in order to integrate inter and intra progeny variability through progeny-specific parameters. The model thus allowed simulating the specificity of plant architecture for a given progeny while including observed inter-individual variability. The architectural model, parameterized for the different progenies, was then implemented in AMAPstudio to generate 3D mock-ups and estimate light interception efficiency, from individual to stand scales.Significant differences in leaf geometry (petiole length, density of leaflets and rachis curvature) and leaflets morphology (gradients of leaflets length and width) were detected between and within progenies, and were accurately simulated by the modelling approach. Besides, light interception estimated from the validated 3D mock-ups showed significant variations among the five progenies.Sensitivity analyses were then performed on a subset of architectural parameters to identify the architectural traits impacting on light interception efficiency and potential carbon assimilation over plant development. The most sensitive parameters over plant development were those related to leaf area (rachis length, number of leaflets, leaflets morphology), but fine attribute related to leaf geometry showed increasing influence when canopy got closed. In adult stand, optimized carbon assimilation was estimated on plants presenting a leaf area index (LAI) between 3.2 and 5.5 m2.m−2, with erected leaves, short rachis and petiole and high number of leaflet on rachis. Four architectural ideotypes for carbon assimilation were proposed based on specific combinations of organs geometry, limiting mutual shading and optimizing light distribution within plant crown.In conclusion, this study highlighted how a functional-structural plant model (FSPM) can be used to virtually explore plant biology. In our case of study, the 3D model of oil palm, in its conception and its application, permitted to detect the architectural traits genetically determined and influencing light interception. The limited number of traits revealed in the sensitivity analysis and the combination of traits proposed through ideotypes could guide further breeding programs. Forthcoming work will be dedicated to integrate in the modeling approach other physiological processes such as stomatal conductance and carbon partitioning. The improved FSPM could then be used to test different scenarios, for instance in climate change context with low radiations or frequent drought events. Similarly, the model could be used to investigate different planting patterns and intercropping systems, and proposed new multi-criteria ideotypes of oil palm

Dans les systèmes agroforestiers simples à deux strates, la distribution de la lumière sous les couronnes des arbres est un facteur clé pour le développement de la culture associée. Cette distribution évolue en fonction des saisons et de la dynamique de croissance de la strate arborée. La lumière disponible pour la culture dépend de la quantité de lumière interceptée par les couronnes des arbres et par conséquent de l'architecture des espèces utilisées. L'objectif de cette thèse est de déterminer le rôle de l'architecture de deux espèces, Acacia mangium Willd. et Tectona grandis L., dans (i) les modalités de l'interception de la lumière par les couronnes et par les feuilles et (ii) les variations intra- et interspécifiques de la distribution du rayonnement sous les couronnes de ces arbres selon leur âge. A partir de l'analyse dynamique des processus de croissance et de ramification de ces deux espèces, un échantillonnage des organes foliaires a permis de déterminer leur répartition, leur géométrie et leurs caractéristiques morphologique, anatomique, optique et biochimique dans les couronnes. Par la simulation de la croissance, de ces deux espèces, paramétrée via des lois mathématiques de croissance et de ramification, des reconstructions 3D d'arbres ont été réalisées en vue de simuler le bilan radiatif de sous-parcelles agroforestières. Les résultats montrent de fortes variations inter- et intraspécifiques de la distribution des organes foliaires et leurs caractéristiques au sein de la couronne et de la distribution du rayonnement sous les couronnes. Les stratégies adoptées par ces deux espèces pour optimiser l'interception lumineuse sont discutées

O presente estudo pretende contribuir para a avaliação da influência de diferentes níveis de adubação azotada nas características agrometeorológicas de desenvolvimento e crescimento do sorgo forrageiro de regadio, nas condições edafo-climáticas do Nordeste Transmontano. Analisou-se o efeito de quatro níveis de azoto no solo na forma como o crescimento e desenvolvimento do sorgo respondem à temperatura e à radiação solar, nomeadamente em termos de crescimento em altura, evolução do número de folhas, índice de área foliar, produção de biomassa, e extracção de azoto. Esta avaliação é feita em termos dos tempos térmicos de desenvolvimento, da eficiência de utilização da radiação para a produção de matéria seca e da eficiência de utilização do azoto. O estudo decorreu na Quinta de Sta. Apolónia, em Bragança, durante 1997, 1998 e 1999, com a variedade de sorgo híbrido forrageiro Sweet Sioux IV. A aplicação de azoto aumentou as áreas foliares, a intercepção da radiação e a produção de biomassa. Contudo, aplicações de N superiores a 400 kg ha-1 não devem ser ultrapassados porque não aportam um aumento significativo de produtividade nem um aumento importante de eficiência de utilização da radiação solar, e reduzem as eficiências de recuperação e de utilização do azoto. The aim of this work is to evaluate the influence of distinct levels of soil nitrogen fertilization on the agrometeorological characteristics of development and growth of irrigated fodder sorghum in the Nordeste Transmontano (Northeastern Portugal). The effects of four levels of soil nitrogen fertilization on the responses of sorghum development and to temperature and solar radiation were analysed, namely crop height, leaf number, leaf area, biomass and nitrogen uptake. This assessment is based on the thermal time for development, on the radiation use efficiency for dry mater production and on the efficiency of nitrogen utilization. The experiments were carried out in Quinta de Sta. Apolónia, Bragança, in 1997, 1998 and 1999, with the hybrid fodder sorghum variety Sweet Sioux IV. Nitrogen fertilization increased leaf area, radiation interception and biomass production. However, nitrogen fertilization rates above 400 kg ha-1 should not be used because they do not lead to a significant increases of both yield and radiation use efficiency, and they reduce the efficiencies of nitrogen recovery and utilization.

Dissecting the soybean grain yield (GY) to approach it as a sum of its associated processes seems a viable approach to explore this trait considering its complex multigenic nature. Monteith (1972, 1977) first defined potential yield as the result of three physiological efficiencies: light interception (Ei), radiation use efficiency (RUE) and harvest index (HI). Though this rationality is not recent, few works assessing these three efficiencies as strategies to improve crops have been carried out. This thesis approaches yield from the perspective of Ei, RUE, and HI to better understand yield as the result of genetic and physiological processes. This study reveals the phenotypic variation, heritability, genetic architecture, and genetic relationships for Ei, RUE, and HI and their relationships with GY and other physiological and phenological variables. Similarly, genomic prediction is presented as a viable strategy to partially overcome the tedious phenotyping of these traits. A large panel of 383 soybean recombinant inbred lines (RIL) with significant yield variation but shrinkage maturity was evaluated in three field environments. Ground measurements of dry matter, photosynthesis (A), transpiration (E), water use efficiency (WUE), stomatal conductance (gs), leaf area index (LAI) and phenology (R1, R5, R8) were measured. Likewise, RGB imagery from an unmanned aircraft system (UAS) were collected with high frequency (~12 days) to estimate the canopy dynamic through the canopy coverage (CC). Light interception was modeled through a logistic curve using CC as a proxy and later compared with the seasonal cumulative solar radiation collected from weather stations to calculate Ei. The total above ground biomass collected during the growing season and its respective cumulative light intercepted were used to derive RUE through linear models fitting, while apparent HI was calculated through the ratio seeds dry matter vs total above-ground dry matter. Additive-genetic correlations, genome wide association (GWA) and whole genome regressions (WGR) were performed to determine the relationship between traits, their association with genomic regions, and the feasibility of predicting these efficiencies through genomic information. Our results revealed moderate to high phenotypic variation for Ei, RUE, and HI. Additive-genetic correlation showed a strong relationship of GY with HI and moderate with RUE and Ei when the whole data set was considered, but negligible contribution of HI on GY when just the top 100 yielding RILs were analyzed. High genetic correlation to grain yield (GY) was also observed for A (0.87) and E (0.67), suggesting increase in GY can be achieved through the improvement of A or E. The GWA analyses showed that Ei is associated with three SNPs; two of them located on chromosome 7 and one on chromosome 11 with no previous quantitative trait loci (QTLs) reported for these regions. RUE is associated with four SNPs on chromosomes 1, 7, 11, and 18. Some of these QTLs are novel, while others are previously documented for plant architecture and chlorophyll content. Two SNPs positioned on chromosome 13 and 15 with previous QTLs reported for plant height and seed set, weight and abortion were associated with HI. WGR showed high predictive ability for Ei, RUE, and HI with maximum correlation ranging between 0.75 to 0.80. Both directed and undirected multivariate explanatory models indicate that HI has a strong relationship with A, average growth rate of canopy coverage for the first 40 days after planting (AGR40), seed-filling (SFL), and reproductive length (RL). According to the path analysis, increase in one standard unit of HI promotes changes in 0.5 standard units of GY, while changes in the same standard unit of RUE, and Ei produce increases on GY of 0.20 and 0.19 standard units. This study presents novel genetic knowledge for Ei, RUE, HI and GY along with a set of tools that may contribute to the development of new cultivars with enhanced light interception, light conversion and optimized dry matter partitioning in soybean. This work not only complements the physiological knowledge already available with the genetic control of traits directly associated with yield, but also represents a pioneer attempt to integrate traditional physiological traits into the breeding process in the context of physiological breeding