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1

Faúndez, Eduardo y Mauricio Silvera. "Sobre la presencia de la chinche de las coníferas occidental Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae) en Uruguay". REVISTA CHILENA DE ENTOMOLOGÍA 45, n.º 4 (16 de octubre de 2019): 549–51. http://dx.doi.org/10.35249/rche.45.4.19.07.

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2

van der Heyden, Torsten. "Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae: Coreinae: Anisoscelini)". REVISTA CHILENA DE ENTOMOLOGÍA 45, n.º 3 (16 de agosto de 2019): 435–37. http://dx.doi.org/10.35249/rche.45.3.19.17.

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3

Kulijer, Dejan. "LEPTOGLOSSUS OCCIDENTALIS (HETEROPTERA: COREIDAE) AND HARMONIA AXYRIDIS (COLEOPTERA: COCCINELLIDAE), TWO NEW INVASIVE ALIEN SPECIES FOR INSECT FAUNA OF MACEDONIA". Ecologica Montenegrina 5 (18 de enero de 2016): 22–25. http://dx.doi.org/10.37828/em.2016.5.3.

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4

Garcete-Barrett, Bolívar R., Sergio D. Rios y Sergio Galeano. "First report of the western conifer seed bug, Leptoglossus occidentalis Heidemann, 1910 (Hemiptera, Coreidae) in Paraguay". Paraquaria Natural 8, n.º 2 (15 de octubre de 2021): 12–14. http://dx.doi.org/10.32525/paraquarianat.2020.(8):12.14.

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5

Strong, Ward B., Sarah L. Bates y Michael U. Stoehr. "Feeding by Leptoglossus occidentalis (Hemiptera: Coreidae) reduces seed set in lodgepole pine (Pinaceae)". Canadian Entomologist 133, n.º 6 (diciembre de 2001): 857–65. http://dx.doi.org/10.4039/ent133857-6.

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AbstractLow seed set is a serious problem in seed orchards of lodgepole pine (Pinus contorta var. latifolia Engelmann) in the southern interior of British Columbia. We tested the hypothesis that Leptoglossus occidentalis Heidemann is responsible for the low seed set. Cones enclosed in insect exclusion bags as part of a pollination experiment produced significantly more filled seeds per cone than cones that were not bagged. In a separate bagging experiment, cones that were enclosed with a L. occidentalis female and her progeny produced only about one filled seed per cone, compared with about 28 seeds in bagged control cones. Changes in microclimate associated with the use of bags did not appear to be responsible for the observed increase in seed set in bagged cones. Leptoglossus occidentalis was also excluded from trees using the insecticide fenvalerate. Cones on fenvalerate-treated trees produced > 11 filled seeds per cone, whereas water-treated (control) cones produced < 1.7 filled seeds. These data suggest that L. occidentalis should be considered a serious pest in lodgepole pine seed orchards.
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6

Kalashian, M. Yu, T. L. Ghrejyan y G. H. Karagyan. "FIRST FINDING OF WESTERN CONIFER SEED BUG LEPTOGLOSSUS OCCIDENTALIS HEID. (HETEROPTERA, COREIDAE) IN ARMENIA". Russian Journal of Biological Invasions 14, n.º 2 (11 de junio de 2021): 52–55. http://dx.doi.org/10.35885/1996-1499-2021-14-2-52-55.

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Data on the first registration of conifer seed bug Leptoglossus occidentalis in Armenia are presented. Penetration of the species from Georgia due to self-dependent expansion or unintentional delivery is presumed.
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7

Lait, Cameron G., Sarah L. Bates, Karen K. Morrissette, John H. Borden y Allison R. Kermode. "Biochemical assays for identifying seeds of lodgepole pine and other conifers fed on by Leptoglossus occidentalis Heidemann (Hemiptera: Coreidae)". Canadian Journal of Botany 79, n.º 11 (1 de noviembre de 2001): 1349–57. http://dx.doi.org/10.1139/b01-119.

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Radiography is a valuable tool for assessing quality of conifer seeds, but it cannot differentiate between aborted seeds and seeds that have been emptied by western conifer seed bug (Leptoglossus occidentalis Heidemann) feeding. We tested three biochemical marker-based assays that were developed to identify L. occidentalis damage to seeds of Douglas-fir, Pseudostuga menziesii (Mirb.) Franco, for their use in lodgepole pine, Pinus contorta var. latifolia Engelmann. The three assays included measurement of storage protein reserve depletion, immunodetection of fragments of insoluble (crystalloid) storage proteins, and immunodetection of L. occidentalis salivary proteins. Aborted seeds contained significantly less soluble and insoluble protein than seeds that were fed on by L. occidentalis. Polyclonal antibodies raised against 11S globulin crystalloid proteins or L. occidentalis salivary gland extracts only immunoreacted with proteins in seeds exposed to L. occidentalis feeding. In a single-blind test, antibody raised against salivary-gland extracts correctly distinguished between undamaged full seeds, unfilled aborted seeds, and seeds fed on by L. occidentalis. Immunodetection of L. occidentalis salivary proteins was also performed on seeds of Abies amabilis Dougl. ex J. Forbes, Tsuga heterophylla (Raf.) Sarg., Picea sitchensis Bong (Carr.), Pinus ponderosa Lawson, and Pinus monticola Dougl. ex D. Don. For all species, immunoreactive polypeptides were only detected in seeds fed on by L. occidentalis. These biochemical marker-based techniques could help researchers and seed orchard managers estimate seed losses caused by L. occidentalis in commercial seed orchards and natural forest stands.Key words: Leptoglossus occidentalis, saliva, biochemical markers, polyclonal antibody, immunodetection, Pinus contorta.
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8

GOMEZ, Cecilia A. "New records of Leptoglossus occidentalis Heidemann (Hemiptera: Coreidae) in Argentine Patagonia". Revista de la Sociedad Entomológica Argentina 80, n.º 04 (30 de diciembre de 2021): 129–32. http://dx.doi.org/10.25085/rsea.800408.

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New records of Leptoglossus occidentalis Heidemann from Andean region is given, becoming the first occurrence within the province of Chubut. The specimens were collected in different environments of urban and periurban areas -both domiciliary and peridomiciliary-, from Esquel, Trevelin and Aldea Escolar. This report extends its southern distribution area to the parallel 43°S.
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9

Petrakis, P. V. "First record of Leptoglossus occidentalis (Heteroptera: Coreidae) in Greece". ENTOMOLOGIA HELLENICA 20, n.º 2 (1 de junio de 2017): 83. http://dx.doi.org/10.12681/eh.11512.

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The Nearctic coreid Leptoglossus occidentalis Heidemann 1910 (Hemiptera: Coreidae) known as the Western Conifer Seed Bug, is recorded for the first time in Greece. This highly dispersible species is well established in Europe and was collected by the authors in various areas of continental Greece (Central Evia, Attica and North Peloponnisos), suggesting a wide distribution in this country.
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10

Campbell, Bruce C. y Patrick J. Shea. "A SIMPLE STAINING TECHNIQUE FOR ASSESSING FEEDING DAMAGE BY LEPTOGLOSSUS OCCIDENTALIS HEIDEMANN (HEMIPTERA: COREIDAE) ON CONES". Canadian Entomologist 122, n.º 5 (octubre de 1990): 963–68. http://dx.doi.org/10.4039/ent122963-9.

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AbstractPectinmethylesterase (PME) activity was found in the salivary glands of nymphs and adults of a leaf-footed bug, Leptoglossus occidentalis Heidemann. Puncture wounds in cone scales resulting from PME activity in the saliva of these bugs were seen by staining with a 0.05% aqueous solution of ruthenium red. This staining technique can be used to estimate feeding damage by L. occidentalis on cones of western white pine (Pinus monticola Dougl.), sugar pine (P. lambertiana Dougl.), and Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco).
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11

van der Heyden, Torsten. "First records of Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae: Coreinae: Anisoscelini) in Finland". REVISTA CHILENA DE ENTOMOLOGÍA 46, n.º 1 (31 de enero de 2020): 73–74. http://dx.doi.org/10.35249/rche.46.1.20.09.

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12

Blatt, Suzie E. y John H. Borden. "EVIDENCE FOR A MALE-PRODUCED AGGREGATION PHEROMONE IN THE WESTERN CONIFER SEED BUG, LEPTOGLOSSUS OCCIDENTALIS HEIDEMANN (HEMIPTERA: COREIDAE)". Canadian Entomologist 128, n.º 4 (agosto de 1996): 777–78. http://dx.doi.org/10.4039/ent128777-4.

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The western conifer seed bug, Leptoglossus occidentalis Heidemann, and the southern pine seed bug, L. corculus (Say), feed on the developing seeds of conifers causing great losses to seed crops (Koerber 1963; Connelly and Schowalter 1991; Schowalter 1996). When high populations threaten the yieldof genetically superior, high quality seed in seed orchards, chemical controls may be used (DeBarr et al. 1982). However, because even minor uses of chemical pesticides are increasingly restricted, particularly in British Columbia where seed orchards commonly interface with urban areas, damaging populations of L. occidentalis may not be controlled at all.
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13

Barta, Marek. "Pathogenicity assessment of entomopathogenic fungi infecting Leptoglossus occidentalis (Heteroptera: Coreidae)." Czech Mycology 62, n.º 1 (15 de febrero de 2010): 67–78. http://dx.doi.org/10.33585/cmy.62107.

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14

Blatt, Suzie E. y John H. Borden. "DISTRIBUTION AND IMPACT OF LEPTOGLOSSUS OCCIDENTALS HEIDEMANN (HEMIPTERA: COREIDAE) IN SEED ORCHARDS IN BRITISH COLUMBIA". Canadian Entomologist 128, n.º 6 (diciembre de 1996): 1065–76. http://dx.doi.org/10.4039/ent1281065-6.

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AbstractLeptoglossus occidentalis Heidemann (Hemiptera: Coreidae) is considered to be an important cone and seed pest in seed orchards in British Columbia; however, no impact studies have been done in Canada. Surveys to determine the distribution of L. occidentalis were conducted over a 3-year period in three coastal seed orchards of Douglas-fir, Pseudotsuga menziesii (Mirb.) Franco, and over a 2-year period in two seed orchards of lodgepole pine, Pinus contorta var. latifolia Engelmann. Leptoglossus occidentalis was found to distribute itself in patches, coinciding with a clonal preference. Seed losses were assessed for 2 years in the coastal Douglas-fir orchards and for 1 year in one lodgepole pine orchard. The percentages of partially filled seed that could have been caused by feeding of L. occidentalis or by a number of other factors were < 5% in coastal Douglas-fir for both 1992 and 1993 and ~14% in lodgepole pine in 1993. Apparently because populations of L. occidentalis were low, there was no correlation between the percentage of partially filled seed and number of insects per tree. Except in unusual circumstances, management measures in Canada are judged not to be necessary as populations of L. occidentalis are consistently below the calculated threshold levels required to cause ≥5% loss in seed production.
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15

Yoon, Chun-Sik, Hyoung-Gon Kim, Jong-Dae Park, Won-Young Choi, Hyeok-Jae Choi y Seon-Woo Cheong. "First Record of the Western Conifer Seed Bug, Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae) in Korea". Journal of Environmental Science International 21, n.º 8 (31 de agosto de 2012): 1009–13. http://dx.doi.org/10.5322/jes.2012.21.8.1009.

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16

Kornev, I., E. Aksenenko y N. Kharchenko. "Invasion Leptoglossus occidentalis Heidemann, 1910 (Heteroptera: Coreidae) in the conditions of Voronezh (Russia)". IOP Conference Series: Earth and Environmental Science 875, n.º 1 (1 de octubre de 2021): 012018. http://dx.doi.org/10.1088/1755-1315/875/1/012018.

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Abstract The article presents new data on the distribution and acclimatization of an alien species of the American coniferous bug (Leptoglossus occidentalis Heidemann, 1910) from the European part of Russia. This bug was discovered in the autumn of 2019 on the territory of the city of Voronezh in the Arboretum of the Voronezh State University of Forestry and Technologies. Currently, L. occidentalis spreads on the territory of Russia only in an urban environment, where it finds optimal environmental conditions (suitable temperature, humidity and shelter for winter locations). Surveys of a number of park and forest park zones, squares and separately growing coniferous trees conducted in 2019-2020 in Voronezh did not reveal the presence of this species. At the moment, in the conditions of the Voronezh region, the center of the spread of this bug is located only in the Arboretum of the Voronezh State University of Forestry and Technologies. In 2020, an increase in the number of the identified group of L. occidentalis was recorded. Further spread of the bug is predicted on the territory of Voronezh. At the moment, no danger to conifers in the European part of Russia from L. occidentalis has been identified and is not yet expected, but constant monitoring of the detected group is required.
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17

Guérin, Bernard. "Leptoglossus occidentals (Heteroptera) : contribution à l'étude de sa progression en France". Bulletin mensuel de la Société linnéenne de Lyon 77, n.º 5 (2008): 65–66. http://dx.doi.org/10.3406/linly.2008.13679.

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18

Lesieur, V., E. Lombaert, T. Guillemaud, B. Courtial, W. Strong, A. Roques y M. A. Auger-Rozenberg. "The rapid spread of Leptoglossus occidentalis in Europe: a bridgehead invasion". Journal of Pest Science 92, n.º 1 (25 de mayo de 2018): 189–200. http://dx.doi.org/10.1007/s10340-018-0993-x.

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19

van der Heyden, Torsten. "Summarized data on the European distribution of Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae: Coreinae: Anisoscelini)". REVISTA CHILENA DE ENTOMOLOGÍA 45, n.º 3 (27 de septiembre de 2019): 499–502. http://dx.doi.org/10.35249/rche.45.3.19.24.

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20

Beránek, J. "First records of Leptoglossus occidentalisdemann, 1910 (Heteroptera: Pentatomorpha: Coreidae) in the Czech Republic". Plant Protection Science 43, No. 4 (7 de enero de 2008): 165–68. http://dx.doi.org/10.17221/2242-pps.

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In the autumn of 2006 and summer 2007 the coreid species <I>Leptoglossus occidentalis</I> Heidemann, 1910 was detected in the Czech Republic for the first time as a new alien insect species. It is native to North America, where it causes important seed losses on coniferous trees, especially pines. From Europe it is known since the end of the last century, when it was introduced to Italy. Its repeated finding could indicate a vital population of this species in the Czech Republic, not only introduced individuals.
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21

Faúndez, Eduardo I., Mariom A. Carvajal y Javier Villablanca. "Alien Invasion: The Case of the Western Conifer-Seed Bug (Heteroptera: Coreidae) in Chile, Overreaction, and Misidentifications". Journal of Medical Entomology 57, n.º 1 (5 de agosto de 2019): 297–303. http://dx.doi.org/10.1093/jme/tjz127.

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Abstract Leptoglossus occidentalis (Heidemann) is an invasive species introduced in Europe, Asia, and most recently South America. In the present study, we report the overreaction situation caused by this bug in Chile, as it has been confused with kissing bugs (Reduviidae: Triatominae), Chagas’ disease vectors. During 2018 and first months of 2019, we received 74 alleged cases of kissing bugs to confirm identification. From these, a total of 63 were identified as L. occidentalis, representing a 85% of the total denounces. Additionally, the first bite case in a human is described. The situation caused by L. occidentalis in Chile is discussed, and an illustrated table is provided to correctly identify this species and tell it apart from Triatomines. It is concluded that L. occidentalis is well established in Chile, and it is necessary to educate the population on recognition of the bug and to avoid overreaction as the species can inflict painful bites, but not transmit any disease.
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22

Gapon, D. A., N. V. Busarova y Yu E. Komarov. "New records of the western conifer seed bug Leptoglossus occidentalis (Heteroptera: Coreidae) in Russia and in adjacent territories". Caucasian Entomological Bulletin 12, n.º 2 (2016): 221–22. http://dx.doi.org/10.23885/1814-3326-2016-12-2-221-222.

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23

Cvetkovska-Gjorgievska, Aleksandra, Ivailo Dedov, Slavcho Hristovski, Mario Langourov, Stanislava Lazarevska, Dana Prelik y Nikolay Simov. "NEW RECORDS OF ALLOCHTONOUS, INVASIVE AND PEST INVERTEBRATE SPECIES FROM THE REPUBLIC OF MACEDONIA". Ecologica Montenegrina 20 (23 de febrero de 2019): 56–70. http://dx.doi.org/10.37828/em.2019.20.5.

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In this paper new data on allochtonous and invasive invertebrate species: Arion (Arion) cf. vulgaris Moquin-Tandon 1855, Cornu aspersum (O.F. Muller, 1774), Harmonia axyridis (Pallas, 1773), Acanthoscelides obtectus (Say, 1831), Trogoderma variabile Ballion, 1878, Stegobium paniceum (Linnaeus, 1758), Oxycarenus (s.str.) lavaterae (Fabricius, 1787), Corythucha ciliata (Say, 1832), Leptoglossus occidentalis Heidemann, 1910, Ceroplastes japonicus (Green, 1921), Pseudaulacaspis pentagona (Targioni Tozzetti, 1886) MacGillivray, 1921, Tuta absoluta (Meyrick, 1917) and Frankliniella occidentalis (Pergande, 1895) from Macedonia are presented. The study represents faunistic data synthesis from different parts of Macedonia, based on collecting efforts of several taxonomic experts. The results of the study help to outline the composition of non-native and invasive invertebrate fauna and to narrow gaps concerning the knowledge of their distribution and introduction in Macedonia.
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24

Bates, Sarah L. "Damage to common plumbing materials caused by overwintering Leptoglossus occidentalis (Hemiptera: Coreidae)". Canadian Entomologist 137, n.º 4 (agosto de 2005): 492–96. http://dx.doi.org/10.4039/n05-005.

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The western conifer seed bug, Leptoglossus occidentalis Heidemann (Hemiptera: Coreidae), is a polyphagous pest of coniferous trees (Hedlin et al. 1981). Native to western North America, the seed bug has expanded its range to include eastern Canada and the United States (McPherson et al. 1990; Gall 1992; Marshall 1992; Ridge-O'Connor 2001; Bates 2002) and was recently introduced into Europe (Taylor et al. 2001). Both adults and nymphs feed by inserting their stylets into cones and digesting the contents of developing seeds, and they can cause serious economic losses in high-value seed orchards (Strong et al. 2001; Bates et al. 2002; Bates and Borden 2005).
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25

Langourov, Mario, Nikolay Simov y Stanislav Abadjiev. "The invasion continues: Hierodula tenuidentata (Mantodea), Leptoglossus occidentalis (Hemiptera) and Pelopidas thrax (Lepidoptera) already on the island of Alonnisos". Historia naturalis bulgarica 44, n.º 11 (20 de diciembre de 2022): 137–44. http://dx.doi.org/10.48027/hnb.44.112.

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During a brief visit of Alonnisos Island in 2022 some insect species (mainly butterflies) were observed and photographed. Three of them are new for the island’s fauna - the giant Asian mantis Hierodula tenuidentata Saussure, 1869, western conifer seed bug Leptoglossus occidentalis Heidemann, 1910, millet skipper Pelopidas thrax (Hübner, [1821]). Their establishment on the island is not a surprise, but it well illustrates the expansion of these species in the Mediterranean area. A list of the observed butterflies is also provided.
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26

Farinha, Ana Oliveira, Manuela Branco, Claudine Courtin, Vincent Lesieur, Diego Gallego, Francisco Javier Sanchez-Garcia, Edmundo Sousa, Alain Roques, Marie-Anne Auger-Rozenberg y Audrey Bras. "Another Chapter in the History of the European Invasion by the Western Conifer Seed Bug, Leptoglossus occidentalis: The Iberian Peninsula". Diversity 15, n.º 1 (4 de enero de 2023): 64. http://dx.doi.org/10.3390/d15010064.

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The Western conifer seed bug, Leptoglossus occidentalis, is native to North America and has already been considered a significant pest in several European countries since its first observation in Italy in 1999. In Spain and Portugal, it was recorded for the first time in 2003 and 2010, respectively, and its impact on Stone Pine (Pinus pinea) is of major concern. Before developing control measures for this insect pest, it is paramount to clarify its spatiotemporal dynamics of invasion. Therefore, in this study, we aimed to (a) characterise the genetic structure and diversity and (b) invasion pathways of L. occidentalis populations in the Iberian Peninsula. To do so, specimens of L. occidentalis were collected at fourteen sites widely distributed within the Iberian Peninsula. We used mtDNA sequences of Cytochrome b and eleven microsatellite markers to characterise the genetic diversity and the population structure in the Iberian Peninsula. Our genetic results combined with the observational dates strongly support a stratified expansion of L. occidentalis invasion in the Iberian Peninsula proceeding from multiple introductions, including at least one in Barcelona, one in Valencia, and one in the west coast or in the Southeastern region.
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27

Luchi, N., V. Mancini, M. Feducci, A. Santini y P. Capretti. "Leptoglossus occidentalis and Diplodia pinea: a new insect-fungus association in Mediterranean forests". Forest Pathology 42, n.º 3 (9 de diciembre de 2011): 246–51. http://dx.doi.org/10.1111/j.1439-0329.2011.00750.x.

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28

Bates, Sarah L., John H. Borden, Allison R. Kermode y Robb G. Bennett. "Impact of Leptoglossus occidentalis (Hemiptera: Coreidae) on Douglas-Fir Seed Production". Journal of Economic Entomology 93, n.º 5 (1 de octubre de 2000): 1444–51. http://dx.doi.org/10.1603/0022-0493-93.5.1444.

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29

Richardson, Tamara A., Ward B. Strong, Brian H. Aukema, Stephen Takàcs, Tracy Zahradnik y B. Staffan Lindgren. "Assessment of cues potentially mediating host selection of Leptoglossus occidentalis on Pinus contorta". Agricultural and Forest Entomology 19, n.º 3 (9 de noviembre de 2016): 245–56. http://dx.doi.org/10.1111/afe.12203.

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30

Kim, Dong Eon, Heejo Lee, Mi Jeong Kim, Young Gyu Ban y Dayeong Kim. "Leptoglossus occidentalis (Hemiptera: Coreidae) occurrence, potential habitats, and COI diversity in South Korea". Journal of Asia-Pacific Biodiversity 13, n.º 1 (marzo de 2020): 35–45. http://dx.doi.org/10.1016/j.japb.2019.11.002.

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31

Bella, Salvatore y Antonio Franquinho Aguiar. "Non-native insect pests from the Madeira Archipelago (Portugal): new records and further data (Insecta: Orthoptera; Thysanoptera; Hemiptera; Coleoptera; Diptera; Lepidoptera; Hymenoptera)". Fragmenta Entomologica 52, n.º 2 (15 de noviembre de 2020): 369–76. http://dx.doi.org/10.4081/fe.2020.469.

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In this study, the results of recent surveys on non-native insect pests from the Madeira Archipelago are reported. Overall, 13 non-native species were recorded: Tessellana tessellata (Charpentier) (Orthoptera, Tettigoniidae), Gynaikothrips ficorum (Marchal) (Thysanoptera, Phlaeothripidae), Leptoglossus occidentalis Heidemann (Hemiptera, Coreidae), Macrohomotoma gladiata Kuwayama (Hemiptera, Homotomidae), Platycorypha nigrivirga Burckhardt and Cacopsylla fulguralis (Kuwayama) (Hemiptera, Psyllidae), Greenidea ficicola Takahashi (Hemiptera, Aphididae), Aloephagus myersi Essig (Hemiptera, Pemphigidae), Protopulvinaria pyriformis (Cockerell) (Hemiptera, Coccidae), Rhynchophorus ferrugineus Olivier (Coleoptera, Dryophthoridae), Phytoliriomyza jacarandae Steyskal & Spencer (Diptera, Agromyzidae), Lantanophaga pusillidactylus (Walker) (Lepidoptera, Pterophoridae), and Josephiella microcarpae Beardsley & Rasplus (Hymenoptera, Agaonidae). Particularly, T. tessellata, L. occidentalis, M. gladiata, P. jacarandae, and J. microcarpae are reported for the first time from the island of Madeira. L. occidentalis is also reported for the first time from the island of Porto Santo. For P. jacarandae this is the first record from Macaronesia, while for M. gladiata this is the first record also for Portugal. Most of these species are associated with allochthonous plants. Details on current distributions, host plants, biological remarks, and natural enemies are given for each species.
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32

Rappaport, N. G., M. I. Haverty, P. J. Shea y R. E. Sandquist. "EFFICACY OF ESFENVALERATE FOR CONTROL OF INSECTS HARMFUL TO SEED PRODUCTION IN DISEASE-RESISTANT WESTERN WHITE PINES". Canadian Entomologist 126, n.º 1 (febrero de 1994): 1–5. http://dx.doi.org/10.4039/ent1261-1.

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AbstractWe tested the pyrethroid insecticide esfenvalerate in single, double, and triple applications for control of insects affecting seed production of blister rust-resistant western white pine, Pinus monticola Douglas. All treatments increased the proportion of normal seed produced and reduced the proportion of seed damaged by the western conifer seed bug, Leptoglossus occidentalis Heidemann. Only the triple application reduced the proportion of cones killed by the pine cone beetle, Conophthorus ponderosae Hopkins. Other seed-damaging insect species [seed chalcids, Megastigmus sp.; the fir coneworm, Dioryctria abietivorella (Grote); and seedworms, Cydia sp.] were present but in numbers too low to test for insecticide efficacy.
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33

Strong, Ward B. "Seasonal changes in seed reduction in lodgepole pine cones caused by feeding of Leptoglossus occidentalis (Hemiptera: Coreidae)". Canadian Entomologist 138, n.º 6 (diciembre de 2006): 888–96. http://dx.doi.org/10.4039/n05-092.

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AbstractManagement of Leptoglossus occidentalis Heidemann in lodgepole pine (Pinus contorta var. latifolia Engelm.) seed orchards of British Columbia, Canada, would be improved with knowledge of its damage potential at different times of the growing season. Mesh insect-exclusion bags were placed over cones, and adults or nymphs of L. occidentalis were enclosed in different bags for 9 periods of 10 to 38 days between 6 May and 17 September 2004. Feeding by adult females between 6 May and 28 May reduced total extractable seeds, a result of ovule damage before fertilization. Reduction in the number of filled seeds per cone was highest between 6 May and 29 June, with each adult female reducing yield by approximately 1.7 seeds per day. Between 29 June and 10 August, each adult female reduced the number of filled seeds per cone (seed set) by 1.0 to 1.25 per day. Seed set reduction declined to approximately 0.25 filled seeds per adult female per day after 10 August. Third to fifth instars caused seed set reduction between 0.6 and 1.2 filled seeds per cone per nymph feeding day from 29 June to 10 August. Utilizing these findings would improve management of L. occidentalis in a commercial seed orchard setting.
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34

Kulijer, Dejan, Osman Mujezinović, Šemso Šarić, Adi Vesnić, Boris Hrašovec y Mirza Dautbašić. "Leptoglossus occidentalis HEIDEMANN, 1910 (Heteroptera: Coreidae) u Bosni i Hercegovini – recentna rasprostranjenost i prvi dokumentirani nalazi". Šumarski list 141, n.º 11-12 (4 de diciembre de 2017): 581–82. http://dx.doi.org/10.31298/sl.141.11-12.5.

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Sjeverno-američka stjenica, Leptoglossus occidentalis Heidemann, 1910, jedna je od 16 neeuropskih vrsta stjenica do danas unešenih u Europu, od kojih 10 potječe upravo iz Sjeverne Amerike (Rabitsch, 2010). Zbog svojeg prirodnog područja pridolaska u Sjevernoj Americi, područja Pacifičke obale omeđena lancem Stjenjaka na zapadu, Amerikanci su je nazvali „zapadnom stjenicom sjemena četinjača” (western conifer seed bug), čime su, uz područja pridolaska, naznačili i njenu štetnost za sjeme drvenastih vrsta iz porodice Pinaceae. Zbog osobite morfološke značajke, spljoštenog proširenja goljenice stražnjih nogu, također je poznata i kao „stjenica listolikog stopala“ (leaf-footed bug). Slovenski autori iskoristili su ovaj naziv kao predložak i dodali mu češer kao objekt prehrane, pa su je nazvali „storževa listonoška“ (Jurc & Jurc, 2005). U svakom slučaju, ova se stjenica nakon dolaska na europski kontinent u Italiji 1999. godine (Taylor et al. 2001.) vrlo brzo širila Europom, pa je u nepunih 10 godina prodrla u velik broj europskih zemalja, od Velike Britanije na zapadu, do Ukrajine i Rusije na istoku, od Norveške na sjeveru do Turske na jugu (Malumphy & Reid, 2007; Fent & Kment, 2011; Gapon, 2013). Među zemljama juga Europe, kojima se proširio ovaj novi invazivni štetnik, našla se i Bosna i Hercegovina. U radu su prikazani svi provjereni i dokumentirani nalazi L. occidentalis na području BiH, od prvog pronalaska i do sada neobjavljenog nalaza iz travnja 2008. godine (zgrada Zemaljskog muzeja BiH u Sarajevu), pa do najsvježijih nalaza ljeti i ujesen 2016. godine (Igman, Neum, Banja Luka). Recentno područje na kojemu je evidentirana ova nova invazivna stjenica u BiH, pregledno je prikazano prostorno označenim podacima nalaza na slici 1. Iako očekivan, značajan je i prvi nalaz ove nove invazivne stjenice u reliktnim sastojinama bora munjike (Pinus heldreichii H. Christ) u masivu Prenja u studenom 2010. godine.Uvid u kronološki slijed i prostornu distribuciju nalaza L. occidentalis u Bosni i Hercegovini od 2008. do 2016. godine ukazuju na razmjerno brzo širenje ovog štetnika i vjerojatno lokalno povećanje populacije, što je zasigurno rezultiralo i njenom lakšom detekcijom. Nalaze se ipak treba shvatiti u kontekstu u kojem su nastali, a to su u velikoj većini bili neciljani, slučajni nalazi entomologa tijekom nekih drugih terenskih istraživanja. Radi uočljivosti i krupnoće same stjenice i činjenice da se ujesen zavlači u ljudske nastambe, nije rijetkost da se često i građanstvo pojavljuje kao „dojavljivač“ novih invazivnih vrsta. Važno je stoga ukazati na nepostojanje sustavnog praćenja pojave i širenja L. occidentalis na ovim prostorima. Uspostava ciljanog monitoringa i praćenja pojave, a posebice potencijalnog štetnog utjecaja L. occidentalis u borovim sastojinama i sjemenskim objektima u Bosni i Hercegovini, nužan su preduvjet potrajnog gospodarenja šumskim bogatstvom i kvalitetniju organizaciju i učinkovitost sustava zaštite šuma od ovog novog invazivnog štetnika.
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35

Lait, Cameron G., Daniel R. Miller, Sarah L. Bates, John H. Borden y Allison R. Kermode. "Biochemical Assay Detects Feeding Damage to Loblolly Pine Seeds Caused by the Leaffooted Pine Seed Bug (Hemiptera: Coreidae)". Journal of Entomological Science 38, n.º 4 (1 de octubre de 2003): 644–53. http://dx.doi.org/10.18474/0749-8004-38.4.644.

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A large number of proteins in salivary gland extracts of the leaffooted pine seed bug, Leptoglossus corculus Say, were strongly recognized by a polyclonal antibody-based assay developed for detecting saliva of the western conifer seed bug. Leptoglossus occidentalis Heidemann, in lodgepole pine, Pinus contorta var. latifolia Engelmann, seeds. An average of approximately 85% of loblolly pine, Pinus taeda L., seeds exposed to feeding by L. corculus for 1 to 4 weeks in the laboratory contained detectable amounts of salivary proteins when the antibody assays were performed weekly on samples (n = 10) of seed. In comparison, radiography of exposed seed detected an average of approximately 63% damaged seed over the same 4-wk period, indicating that the antibody assay increased sensitivity of damage detection by approximately one-third. Depletion of insoluble polypeptides and proliferation of soluble polypeptides ≤23.5 kDa was apparent after SDS-PAGE and quantitative assays were performed on proteins extracted from seeds that were damaged by exposure to L. corculus feeding. Our data suggest that the antibody-based test could be used to obtain accurate estimates of seed losses attributable to L. corculus feeding in southern pine seed orchards.
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36

Taszakowski, Artur, Adrian Masłowski y Jolanta Brożek. "Labial Sensory Organs of Two Leptoglossus Species (Hemiptera: Coreidae): Their Morphology and Supposed Function". Insects 14, n.º 1 (28 de diciembre de 2022): 30. http://dx.doi.org/10.3390/insects14010030.

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A detailed description of the labial sensory organs of Leptoglossus occidentalis Heidemann, 1910 and L. zonatus (Dallas, 1852) (Hemiptera: Heteroptera: Coreidae) is presented. The detailed morphology, location, and distribution of different sensilla types on mouthparts were investigated and shown in micrographs taken with a scanning electron microscope. Nine types of aporous sensilla, and three uniporous sensilla were found. The possible functions of these sensilla as well as similarities and differences between the mouthparts of Leptoglossus and those of other terrestrial Heteroptera are discussed. The tip of the labium constitutes a functional “touch and taste area”, combining the chemosensitivity of uniporous sensilla P1–P3 and the mechanoreceptivity of A8 and A9 hair-like sensilla. A set of two cone-like chemosensilla types (9 + 2) was found on each lateral lobe of the labial tip. Literature analysis showed that such a set of cone-like sensory organs on the labial tip may be common in terrestrial Heteroptera. This observation confirms that the number and arrangement of sensilla is conservative and can be important in diagnosing taxa at various levels and in phylogenetic studies based on morphology.
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37

El Khoury, Yara, Elise Noujeim, Giovanni Bubici, Eustachio Tarasco, Charbel Al Khoury y Nabil Nemer. "Potential Factors behind the Decline of Pinus pinea Nut Production in Mediterranean Pine Forests". Forests 12, n.º 9 (28 de agosto de 2021): 1167. http://dx.doi.org/10.3390/f12091167.

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Mediterranean stone pine nut is appreciated for its high economic and nutritional value. Starting in 2012, Pinus pinea nut production declined throughout the Mediterranean area. The dry cone syndrome associated with this decline and the introduction of Leptoglossus occidentalis occurred simultaneously. This study aims to evaluate potential reasons behind the decline in pine nut production in Lebanon, considering climatic factors and the invasion of L. occidentalis. Correlation analysis was used to examine a potential relationship between cone yield and the percentage of damaged seeds per cone. Climatic variables were also tested. Two time periods were considered for analysis: before and after 2012. Cone production and the percentage of damaged seeds were negatively correlated (r = −0.42). From 2012 to 2017, cone production declined by 50% and the percentage of damaged seeds increased on average from 3% in 2012 up to 60% in 2017. Correlations were detected between cone production and the temperature of the hottest three months of the year of harvesting, and between cone production and average temperatures during the year of cone initiation. A conjunction of factors that include L. occidentalis and climatic factors might have affected the pine nut production in Lebanon.
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38

Farinha, Ana O., C. Carvalho, A. C. Correia y M. Branco. "Impact assessment of Leptoglossus occidentalis in Pinus pinea: Integrating population density and seed loss". Forest Ecology and Management 496 (septiembre de 2021): 119422. http://dx.doi.org/10.1016/j.foreco.2021.119422.

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39

Loewe-Muñoz, Verónica, Claudia Delard, Rodrigo Del Río y Mónica Balzarini. "Western conifer seed bug (Leptoglossus occidentalis) challenging stone pine cropping in the Southern Hemisphere". Forest Ecology and Management 496 (septiembre de 2021): 119434. http://dx.doi.org/10.1016/j.foreco.2021.119434.

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40

Kalashian, M. Yu, T. L. Ghrejyan y G. H. Karagyan. "First Finding of Western Conifer Seed Bug Leptoglossus occidentalis Heid. (Heteroptera, Coreidae) in Armenia". Russian Journal of Biological Invasions 12, n.º 3 (julio de 2021): 274–76. http://dx.doi.org/10.1134/s2075111721030073.

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41

Arslangündoğdu, Zeynel y Erdem Hızal. "The Western Conifer Seed Bug,Leptoglossus occidentalis(Heidemann, 1910), recorded in Turkey (Heteroptera: Coreidae)". Zoology in the Middle East 50, n.º 1 (enero de 2010): 138–39. http://dx.doi.org/10.1080/09397140.2010.10638426.

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42

Bates, Sarah L., John H. Borden, Anne Savoie, Suzanne E. Blatt, Cameron G. Lait, Allison R. Kermode y Robb G. Bennett. "IMPACT OF FEEDING BY LEPTOGLOSSUS OCCIDENTALS (HEMIPTERA: COREIDAE) ON THE MAJOR STORAGE RESERVES OF MATURE DOUGLAS-FIR (PINACEAE) SEEDS". Canadian Entomologist 132, n.º 1 (febrero de 2000): 91–102. http://dx.doi.org/10.4039/ent13291-1.

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AbstractIn laboratory experiments adults and nymphs of the western conifer seed bug, Leptoglossus occidentalis Heidemann, were allowed to feed on mature seeds of Douglas-fir, Pseudotsuga menziesii (Mirbel) Franco. Weight-loss measurements and scanning electron microscopy provided strong supporting evidence for the use of simple radiographic diagnosis as a method of classifying feeding damage to seeds into four categories: light (greater than two thirds of seed contents remaining), moderate (one third to two thirds of seed contents remaining), severe (less than one third of seed contents remaining), and extreme (seed empty). Scanning electron micrographs showed the apparent depletion of lipid and protein storage reserves which was verified by quantitative analyses that showed significant loss of lipid and buffer-insoluble (crystalloid) storage protein from seeds in all damage categories. The amount of buffer-soluble (matrix) protein was reduced in seeds from the severe and extreme damage categories. The increase in buffer-soluble protein observed in lightly damaged seeds was likely due to the solubilization of crystalloid storage protein, as a result of its breakdown into smaller peptides. Our results suggest that through the action of both lipases and proteases, L. occidentalis can have a serious impact on the major storage reserves of conifer seeds. Moreover, our data suggest that L. occidentalis feeds in a different manner than the laceration and flushing method found in other seed-feeding Hemiptera.
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43

Takács, Stephen, Hannah Bottomley, Iisak Andreller, Tracy Zaradnik, Joseph Schwarz, Robb Bennett, Ward Strong y Gerhard Gries. "Infrared radiation from hot cones on cool conifers attracts seed-feeding insects". Proceedings of the Royal Society B: Biological Sciences 276, n.º 1657 (21 de octubre de 2008): 649–55. http://dx.doi.org/10.1098/rspb.2008.0742.

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Foraging animals use diverse cues to locate resources. Common foraging cues have visual, auditory, olfactory, tactile or gustatory characteristics. Here, we show a foraging herbivore using infrared (IR) radiation from living plants as a host-finding cue. We present data revealing that (i) conifer cones are warmer and emit more near-, mid- and long-range IR radiation than needles, (ii) cone-feeding western conifer seed bugs, Leptoglossus occidentalis (Hemiptera: Coreidae), possess IR receptive organs and orient towards experimental IR cues, and (iii) occlusion of the insects' IR receptors impairs IR perception. The conifers' cost of attracting cone-feeding insects may be offset by occasional mast seeding resulting in cone crops too large to be effectively exploited by herbivores.
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44

Blatt, S. E. y J. H. Borden. "INTERACTIONS BETWEEN THE DOUGLAS-FIR SEED CHALCID, MEGASTIGMUS SPERMOTROPHUS (HYMENOPTERA: TORYMIDAE), AND THE WESTERN CONIFER SEED BUG, LEPTOGLOSSUS OCCIDENTALIS (HEMIPTERA: COREIDAE)". Canadian Entomologist 130, n.º 6 (diciembre de 1998): 775–82. http://dx.doi.org/10.4039/ent130775-6.

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AbstractIn two laboratory experiments, adult western conifer seed bugs, Leptoglossus occidentalis Heidemann, did not feed on seed infested by the Douglas-fir seed chalcid, Megastigmus spermotrophus Wachtl. When presented with seed lots containing 0, 20, 40, 60, 80, or 100% chalcid-infested seed, seed bugs fed on uninfested seed only when the percent chalcid infestation was ≤ 40% (Exp. 1), ≤ 20% (Exp. 2, females), or ≤ 60% (Exp. 2, males). In a third experiment, nymphs exposed to similar seed lots did not feed on M. spermotrophus when sound seeds were present. However, when exposed to 100% chalcid-infested seed, nymphs in two of 10 replicates fed on M. spermotrophus. Feeding by seed bugs caused seeds to lose 51% of their weight on average, but 18% of 43 test seeds on which seed bugs had fed germinated. In three Douglas-fir orchards surveyed, both species exhibited a clonal preference, but the ranking of clones preferred by each species was not the same. These results suggest that the impacts of L. occidentalis and M. spermotrophus are segregated and additive.
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45

Adamska, Iwona y Magdalena Dzięgielewska. "Can the western conifer seed bug Leptoglossus occidentalis (Heidemann, 1910) threaten coniferous forests in Poland?" Ecological Questions 32, n.º 1 (13 de agosto de 2020): 1. http://dx.doi.org/10.12775/eq.2021.001.

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46

Faúndez, Eduardo I., Mariom Carvajal y Norka Contreras. "New records of the Western Conifer Seed Bug Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae) in Chile". Revista de la Sociedad Entomológica Argentina 78, n.º 3 (26 de septiembre de 2019): 26–28. http://dx.doi.org/10.25085/rsea.780305.

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47

Barta, Marek. "Biology and temperature requirements of the invasive seed bug Leptoglossus occidentalis (Heteroptera: Coreidae) in Europe". Journal of Pest Science 89, n.º 1 (23 de mayo de 2015): 31–44. http://dx.doi.org/10.1007/s10340-015-0673-z.

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48

Farinha, Ana O., Charlene Durpoix, Susana Valente, Edmundo Sousa, Alain Roques y Manuela Branco. "The stone pine, Pinus pinea L., a new highly rewarding host for the invasive Leptoglossus occidentalis". NeoBiota 41 (21 de diciembre de 2018): 1–18. http://dx.doi.org/10.3897/neobiota.41.30041.

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The invasive seed bugLeptoglossusoccidentalis, a species native to Western North America, is of major concern for the producers of stone pine seeds in the Mediterranean countries. The large size of these edible seeds and their nutritive content may represent a pull factor for the seed bug. Cone and seed traits of three main Mediterranean pine species:P.pinea,P.pinaster, andP.halepensis, were evaluated. Preference trials with cone-bearing branches, individual cones and seeds were conducted to test host preference among the three host species.Considering the kernel size, stone pine seeds provide 4 to 13 times more reward thanP.pinasterandP.halepensisseeds, respectively, but also needed a greater effort to be reached as measured by coat thickness. Still, the benefit/cost ratio was higher onP.pinea. Individual seeds and cones ofP.pineawere 2 to 3 times more consumed than those of the two other pine species. However, branch preference trials did not reveal any difference in bug visits. Moreover, adults manifested strong group behaviour on branches, frequently dissociating into two persisting groups. The implications of these results forP.pineaproducing areas are discussed.
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49

Nemer, Nabil, Yara El Khoury, Elise Noujeim, Yara Zgheib, Eustachio Tarasco y Torsten van der Heyden. "First records of the invasive species Leptoglossus occidentalis Heidemann (Hemiptera: Coreidae) on different coniferous species including the cedars of Lebanon". REVISTA CHILENA DE ENTOMOLOGÍA 45, n.º 4 (16 de octubre de 2019): 507–13. http://dx.doi.org/10.35249/rche.45.4.19.01.

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Roversi, P. F., G. Sabbatini Peverieri, M. Maltese, P. Furlan, W. B. Strong y V. Caleca. "Pre-release risk assessment of the egg-parasitoid Gryon pennsylvanicum for classical biological control of Leptoglossus occidentalis". Journal of Applied Entomology 138, n.º 1-2 (15 de junio de 2013): 27–35. http://dx.doi.org/10.1111/jen.12062.

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