Academic literature on the topic 'Theropods'

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Journal articles on the topic "Theropods"

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Rowe, Timothy. "The Early History of Theropods." Short Courses in Paleontology 2 (1989): 100–112. http://dx.doi.org/10.1017/s2475263000000891.

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Theropods have traditionally been portrayed as extinct bipedal predators built along the lines of such celebrated terrors as Tyrannosaurus, Deinonychus, and Allosaurus. The earliest theropods indeed fit that image, and all of them are decidedly extinct. However, it has become increasingly apparent that living birds trace their genealogy to those extinct theropods (Ostrom, 1976; Gauthier, 1986; Gauthier and Padian, 1985 and this volume), and that any adequate consideration of the evolutionary history of Theropoda must assess the lineage as a whole, instead of arbitrarily focusing on the Mesozoic forms alone. When the approximately 8,600 living avian descendants of the ancestral theropod are taken into account, together with the various extinct Mesozoic and Cenozoic taxa, theropods display a far greater range of size, form, behavior, and diet than we ever pictured in our traditional image.
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Button, Khai, Hailu You, James I. Kirkland, and Lindsay Zanno. "Incremental growth of therizinosaurian dental tissues: implications for dietary transitions in Theropoda." PeerJ 5 (December 11, 2017): e4129. http://dx.doi.org/10.7717/peerj.4129.

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Previous investigations document functional and phylogenetic signals in the histology of dinosaur teeth. In particular, incremental lines in dentin have been used to determine tooth growth and replacement rates in several dinosaurian clades. However, to date, few studies have investigated the dental microstructure of theropods in the omnivory/herbivory spectrum. Here we examine dental histology of Therizinosauria, a clade of large-bodied theropods bearing significant morphological evidence for herbivory, by examining the teeth of the early-diverging therizinosaurian Falcarius utahensis, and an isolated tooth referred to Suzhousaurus megatherioides, a highly specialized large-bodied representative. Despite attaining some of the largest body masses among maniraptoran theropod dinosaurs, therizinosaurian teeth are diminutive, measuring no more than 0.90 cm in crown height (CH) and 0.38 cm in crown base length (CBL). Comparisons with other theropods and non-theropodan herbivorous dinosaurs reveals that when controlling for estimated body mass, crown volume in therizinosaurians plots most closely with dinosaurs of similar dietary strategy as opposed to phylogenetic heritage. Analysis of incremental growth lines in dentin, observed in thin sections of therizinosaurian teeth, demonstrates that tooth growth rates fall within the range of other archosaurs, conforming to hypothesized physiological limitations on the production of dental tissues. Despite dietary differences between therizinosaurians and hypercarnivorous theropods, the types of enamel crystallites present and their spatial distribution—i.e., the schmelzmuster of both taxa—is limited to parallel enamel crystallites, the simplest form of enamel and the plesiomorphic condition for Theropoda. This finding supports previous hypotheses that dental microstructure is strongly influenced by phylogeny, yet equally supports suggestions of reduced reliance on oral processing in omnivorous/herbivorous theropods rather than the microstructural specializations to diet exhibited by non-theropodan herbivorous dinosaurs. Finally, although our sample is limited, we document a significant reduction in the rate of enamel apposition contrasted with increased relative enamel thickness between early and later diverging therizinosaurians that coincides with anatomical evidence for increased specializations to herbivory in the clade.
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Van Valkenburgh, Blaire, and Ralph E. Molnar. "Dinosaurian and mammalian predators compared." Paleobiology 28, no. 4 (2002): 527–43. http://dx.doi.org/10.1666/0094-8373(2002)028<0527:dampc>2.0.co;2.

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Theropod dinosaurs were, and mammalian carnivores are, the top predators within their respective communities. Beyond that, they seem distinct, differing markedly in body form and ancestry. Nevertheless, some of the same processes that shape mammalian predators and their communities likely were important to dinosaurian predators as well. To explore this, we compared the predatory adaptations of theropod dinosaurs and mammalian carnivores, focusing primarily on aspects of their feeding morphology (skulls, jaws, and teeth). We also examined suites of sympatric species (i.e., ecological guilds) of predatory theropods and mammals, emphasizing species richness and the distribution of body sizes within guilds. The morphological comparisons indicate reduced trophic diversity among theropods relative to carnivorans, as most or all theropods with teeth appear to have been hypercarnivorous. There are no clear analogs of felids, canids, and hyaenids among theropods. Interestingly, theropods parallel canids more so than felids in cranial proportions, and all theropods appear to have had weaker jaws than carnivorans. Given the apparent trophic similarity of theropods and their large body sizes, it was surprising to find that species richness of theropod guilds was as great as or exceeded that observed among mammalian carnivore guilds. Separation by body size appears to be slightly greater among sympatric theropods than carnivorans, but the magnitude of size difference between species is not constant in either group. We suggest that, as in modern carnivoran guilds, smaller theropod species might have adapted to the threats posed by much larger species (e.g., tyrannosaurs) by hunting in groups, feeding rapidly, and avoiding encounters whenever possible. This would have favored improved hunting skills and associated adaptations such as agility, speed, intelligence, and increased sensory awareness.
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Gatesy, Stephen M. "Caudofemoral musculature and the evolution of theropod locomotion." Paleobiology 16, no. 2 (1990): 170–86. http://dx.doi.org/10.1017/s0094837300009866.

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Living crocodilians and limbed lepidosaurs have a large caudofemoralis longus muscle passing from tail to femur. Anatomical and electromyographic data support the conclusion that the caudofemoralis is the principal femoral retractor and thus serves as the primary propulsive muscle of the hind limb. Osteological evidence of both origin and insertion indicates that a substantial caudofemoralis longus was present in archosaurs primitively and was retained in the clades Dinosauria and Theropoda. Derived theropods (e.g., ornithomimids, deinonychosaurs, Archaeopteryx and birds) exhibit features that indicate a reduction in caudofemoral musculature, including fewer caudal vertebrae, diminished caudal transverse processes, distal specialization of the tail, and loss of the fourth trochanter. This trend culminates in ornithurine birds, which have greatly reduced tails and either have a minute caudofemoralis longus or lack the muscle entirely.As derived theropod dinosaurs, birds represent the best living model for reconstructing extinct nonavian theropods. Bipedal, digitigrade locomotion on fully erect limbs is an avian feature inherited from theropod ancestors. However, the primitive saurian mechanisms of balancing the body (with a large tail) and retracting the limb (with the caudofemoralis longus) were abandoned in the course of avian evolution. This strongly suggests that details of the orientation (subhorizontal femur) and movement (primarily knee flexion) of the hind limb in extant birds are more properly viewed as derived, uniquely avian conditions, rather than as retentions of an ancestral dinosaurian pattern. Although many characters often associated with extant birds appeared much earlier in theropod evolution, reconstructing the locomotion of all theropods as completely birdlike ignores a wealth of differences that characterize birds.
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Griffin, Christopher T. "Large neotheropods from the Upper Triassic of North America and the early evolution of large theropod body sizes." Journal of Paleontology 93, no. 5 (March 26, 2019): 1010–30. http://dx.doi.org/10.1017/jpa.2019.13.

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AbstractLarge body sizes among nonavian theropod dinosaurs is a major feature in the evolution of this clade, with theropods reaching greater sizes than any other terrestrial carnivores. However, the early evolution of large body sizes among theropods is obscured by an incomplete fossil record, with the largest Triassic theropods represented by only a few individuals of uncertain ontogenetic stage. Here I describe two neotheropod specimens from the Upper Triassic Bull Canyon Formation of New Mexico and place them in a broader comparative context of early theropod anatomy. These specimens possess morphologies indicative of ontogenetic immaturity (e.g., absence of femoral bone scars, lack of co-ossification between the astragalus and calcaneum), and phylogenetic analyses recover these specimens as early-diverging neotheropods in a polytomy with other early neotheropods at the base of the clade. Ancestral state reconstruction for body size suggests that the ancestral theropod condition was small (~240 mm femur length), but the ancestral neotheropod was larger (~300–340 mm femur length), with coelophysoids experiencing secondary body size reduction, although this is highly dependent on the phylogenetic position of a few key taxa. Theropods evolved large body sizes before the Triassic–Jurassic extinction, as hypothesized in most other ancestral state reconstructions of theropod body sizes, but remained rare relative to smaller theropods until the Jurassic.
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Hendrickx, Christophe, Ricardo Araújo, and Octávio Mateus. "The non-avian theropod quadrate I: standardized terminology with an overview of the anatomy and function." PeerJ 3 (September 17, 2015): e1245. http://dx.doi.org/10.7717/peerj.1245.

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The quadrate of reptiles and most other tetrapods plays an important morphofunctional role by allowing the articulation of the mandible with the cranium. In Theropoda, the morphology of the quadrate is particularly complex and varies importantly among different clades of non-avian theropods, therefore conferring a strong taxonomic potential. Inconsistencies in the notation and terminology used in discussions of the theropod quadrate anatomy have been noticed, including at least one instance when no less than eight different terms were given to the same structure. A standardized list of terms and notations for each quadrate anatomical entity is proposed here, with the goal of facilitating future descriptions of this important cranial bone. In addition, an overview of the literature on quadrate function and pneumaticity in non-avian theropods is presented, along with a discussion of the inferences that could be made from this research. Specifically, the quadrate of the large majority of non-avian theropods is akinetic but the diagonally oriented intercondylar sulcus of the mandibular articulation allowed both rami of the mandible to move laterally when opening the mouth in many of theropods. Pneumaticity of the quadrate is also present in most averostran clades and the pneumatic chamber—invaded by the quadrate diverticulum of the mandibular arch pneumatic system—was connected to one or several pneumatic foramina on the medial, lateral, posterior, anterior or ventral sides of the quadrate.
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Feduccia, Alan. "FANTASY VS REALITY: A Critique of Smith et al.'s Bird Origins." Open Ornithology Journal 9, no. 1 (April 30, 2016): 14–38. http://dx.doi.org/10.2174/1874453201609010014.

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Adherents of the current orthodoxy of a derivation of birds from theropod dinosaurs, criticize the commentary by Feduccia (2013, Auk, 130) [1 - 12] entitled “Bird Origins Anew” as well as numerous papers by Lingham-Soliar on theropod dermal fibers, using numerous mischaracterizations and misstatements of content, and illustrate their own misconceptions of the nature of the debate, which are here clarified. While there is general agreement with the affinity of birds and maniraptorans, the widely accepted phylogeny, advocating derived earth-bound maniraptorans giving rise to more primitive avians (i.e. Archaeopteryx), may be “topsy-turvy.” The current primary debate concerns whether maniraptorans are ancestral or derived within the phylogeny, and whether many maniraptorans and birds form a clade distinct from true theropods. Corollaries of the current scheme show largely terrestrial maniraptoran theropods similar to the Late CretaceousVelociraptorgiving rise to avians, and flight originatingviaa terrestrial (cursorial) “gravity-resisted,” as opposed to an arboreal “gravity-assisted” model. The current dogma posits pennaceous flight remiges in earth-bound theropods having evolved in terrestrial theropods that never flew. As part of the orthodoxy, fully feathered maniraptorans such as the tetrapteryx glidersMicroraptorand allies, are incorrectly reconstructed as terrestrial cursors, when in reality their anatomy and elongate hindlimb feathers would be a hindrance to terrestrial locomotion.The same is true of many early birds, exemplified by reconstruction of the arboreally adaptedConfuciusornisas a terrestrial predator, part of the overall theropodan scheme of birds evolving from terrestrial dinosaurs, and flight from the ground up. Both sides of this contentious debate must be constantly aware that new fossil or even molecular discoveries on birds may change current conclusions.
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Zelenitsky, Darla K., François Therrien, and Yoshitsugu Kobayashi. "Olfactory acuity in theropods: palaeobiological and evolutionary implications." Proceedings of the Royal Society B: Biological Sciences 276, no. 1657 (October 28, 2008): 667–73. http://dx.doi.org/10.1098/rspb.2008.1075.

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This research presents the first quantitative evaluation of the olfactory acuity in extinct theropod dinosaurs. Olfactory ratios (i.e. the ratio of the greatest diameter of the olfactory bulb to the greatest diameter of the cerebral hemisphere) are analysed in order to infer the olfactory acuity and behavioural traits in theropods, as well as to identify phylogenetic trends in olfaction within Theropoda. A phylogenetically corrected regression of olfactory ratio to body mass reveals that, relative to predicted values, the olfactory bulbs of (i) tyrannosaurids and dromaeosaurids are significantly larger, (ii) ornithomimosaurs and oviraptorids are significantly smaller, and (iii) ceratosaurians, allosauroids, basal tyrannosauroids, troodontids and basal birds are within the 95% CI. Relative to other theropods, olfactory acuity was high in tyrannosaurids and dromaeosaurids and therefore olfaction would have played an important role in their ecology, possibly for activities in low-light conditions, locating food, or for navigation within large home ranges. Olfactory acuity was the lowest in ornithomimosaurs and oviraptorids, suggesting a reduced reliance on olfaction and perhaps an omnivorous diet in these theropods. Phylogenetic trends in olfaction among theropods reveal that olfactory acuity did not decrease in the ancestry of birds, as troodontids, dromaeosaurids and primitive birds possessed typical or high olfactory acuity. Thus, the sense of smell must have remained important in primitive birds and its presumed decrease associated with the increased importance of sight did not occur until later among more derived birds.
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XI-JIN, ZHAO, ROGER B. J. BENSON, STEPHEN L. BRUSATTE, and PHILIP J. CURRIE. "The postcranial skeleton ofMonolophosaurus jiangi(Dinosauria: Theropoda) from the Middle Jurassic of Xinjiang, China, and a review of Middle Jurassic Chinese theropods." Geological Magazine 147, no. 1 (July 9, 2009): 13–27. http://dx.doi.org/10.1017/s0016756809990240.

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AbstractThe Middle Jurassic was a critical time in the evolution of theropod dinosaurs, highlighted by the origination and radiation of the large-bodied and morphologically diverse Tetanurae. Middle Jurassic tetanurans are rare but have been described from Europe, South America and China. In particular, China has yielded a number of potential basal tetanurans, but these have received little detailed treatment in the literature. Here we redescribe the postcranial skeleton of one of the most complete Chinese Middle Jurassic theropods,Monolophosaurus. Several features confirm the tetanuran affinities ofMonolophosaurus, but the possession of ‘primitive’ traits such as a double-faceted pubic peduncle of the ilium and a hood-like supracetabular crest suggest a basal position within Tetanurae. This conflicts with most published cladistic analyses that placeMonolophosaurusin a more derived position within Allosauroidea. We review the Middle Jurassic record of Chinese theropods and compareMonolophosaurusto other Middle Jurassic theropods globally. These comparisons suggest thatMonolophosaurusandChuandongocoelurusformed an endemic theropod clade limited to the Middle Jurassic of Asia. Other Middle Jurassic Chinese theropods deserve further study.
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Lautenschlager, Stephan. "Morphological and functional diversity in therizinosaur claws and the implications for theropod claw evolution." Proceedings of the Royal Society B: Biological Sciences 281, no. 1785 (June 22, 2014): 20140497. http://dx.doi.org/10.1098/rspb.2014.0497.

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Therizinosaurs are a group of herbivorous theropod dinosaurs from the Cretaceous of North America and Asia, best known for their iconically large and elongate manual claws. However, among Therizinosauria, ungual morphology is highly variable, reflecting a general trend found in derived theropod dinosaurs (Maniraptoriformes). A combined approach of shape analysis to characterize changes in manual ungual morphology across theropods and finite-element analysis to assess the biomechanical properties of different ungual shapes in therizinosaurs reveals a functional diversity related to ungual morphology. While some therizinosaur taxa used their claws in a generalist fashion, other taxa were functionally adapted to use the claws as grasping hooks during foraging. Results further indicate that maniraptoriform dinosaurs deviated from the plesiomorphic theropod ungual morphology resulting in increased functional diversity. This trend parallels modifications of the cranial skeleton in derived theropods in response to dietary adaptation, suggesting that dietary diversification was a major driver for morphological and functional disparity in theropod evolution.
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Dissertations / Theses on the topic "Theropods"

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Rauhut, Oliver Walter Mischa. "The interrelationships and evolution of basal theropods (Dinosauria, Saurischia)." Thesis, University of Bristol, 2000. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.310688.

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Signore, Marco. "Scipionyx samniticus (Therpoda, maniraptoriformes) and the palaeobiology of some maniraptoran theropods." Thesis, University of Bristol, 2001. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.368643.

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Brusatte, Stephen L. "The skull of Monolophosaurus jingi (Dinosauria: Theropoda) and its implications for early theropod phylogeny and evolution." Thesis, University of Bristol, 2008. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.492554.

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The Middle Jurassic was a critical time in the evolution of theropod dinosaurs, highlighted by the origination and initial radiation of the large-bodied and morphologically diverse Tetanurae. Middle Jurassic tetanurans are rare but have been described from Europe, South America, and China. In particular, China has yielded a number of potential basal tetanurans but these have received little detailed treatment in the literature.
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Tahara, Rui. "Cranial pneumaticity of «Ornithomimus edmontonicus» (Ornithomimidae: Theropoda)." Thesis, McGill University, 2009. http://digitool.Library.McGill.CA:80/R/?func=dbin-jump-full&object_id=40806.

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Modern archosaurs have extensive pneumatic invasions derived from nasal and tympanic sinuses. These are present in many fossil archosaurs, but their evolutionary history has yet to be clarified. A full description of the cranial pneumaticity of a well-preserved ornithomimid theropod is presented to help clarify the evolution of this soft tissue using CT scan data and 3D reconstruction. The cranial sinuses of Ornithomimus edmontonicus represent nearly all cranial sinuses of birds and add new information to the range of that of ornithomimids. Phylogenetic comparisons of cranial pneumaticity across theropods with emphasis on O. edmontonicus imply a novel homology between the jugal fossa or recess of non-avian theropods and the suborbital diverticulum of birds. Comparisons also establish the presence of an avian-like nasal sinus morphology at Neotetanurae and tympanic sinus morphology at Coelurosauria.
Les archosauriens modernes ont des invasions pneumatiques extensives dérivées de leurs sinus nasales et tympaniques. Ils sont présents dans de nombreux fossiles d’archosauriens, mais l’histoire de leur évolution demeure incertaine. Une description complète de la pneumaticité crâniens d’un théropode ornithomimidé bien préservé aide à clarifier l’évolution de ces tissues mous à l’aide de donnés d’un scanneur CT et de la reconstruction 3D. Les sinus crâniennes de Omithomimus edmontonicus représentent presque tous les sinus crâniens des oiseaux et rajoutent de nouvelles informations en ce qui à trait aux ornithomimidés. Les comparaisons phylogénétiques de la pneumaticité crânienne à travers les théropodes avec une emphase sur O. edmontonicus suggèrent une nouvelle homologie de la jugal fossa ou la dépression des théropodes non-aviaires et le diverticulum sous-orbital des oiseaux. Les comparaisons permettent aussi d’établir la présence de sinus nasals à morphologie aviaire au Neotetanurae et de sinus à morphologie tympanique au Coelurosauria.
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Silva, Rosely Rodrigues da. "Descrição osteológica e posicionamento filogenético de um terópode (Dinosauria, Saurischia) do Cretáceo Inferior da Bacia Sanfranciscana, município de Coração de Jesus, Minas Gerais, Brasil." Universidade de São Paulo, 2013. http://www.teses.usp.br/teses/disponiveis/41/41133/tde-22072013-110420/.

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A região do cráton São Francisco possui um histórico de eventos de sedimentações que datam do Paleo-Proterozóico e Mesozóico. A Bacia Sanfranciscana representa o último ciclo geodinâmico que afetou a grande Bacia Intracratônica do São Francisco. Embora o Cretáceo brasileiro seja considerado o período geológico mais bem representativo em termos de conteúdo de paleofauna de vertebrados, na Bacia Sanfranciscana este registro era restrito até recentemente apenas por celacantídeos e fragmentos de peixes do Grupo Areado. Através dos trabalhos de campos executados pela equipe do Laboratório de Paleontologia do Museu de Zoologia da USP, novos fósseis de tetrápodes continentais foram encontrados em sedimentos da Formação Quiricó, nos arredores do município de Coração de Jesus (MG). Dentre estes materiais, destacam-se elementos epipodiais, mesopodiais e metapodiais atribuíveis a um terópode. Este estudo visou descrever e comparar a osteologia do novo exemplar, tombado sob o número MZSP-PV 830, com outros registros de abelissauroideos conhecidos além de posicionar filogeneticamente com os outros táxons da irradiação de terópodes formada pelo clado Ceratosauria. Neste sentido, a análise da matriz de dados alocou-o no clado Noasauridae, com base nas seguintes características: a forma tabular do maléolo lateral da tíbia; e a forma estreita da haste do metatarsal II. A presente descrição representa uma relevância paleontológica significativa para o entendimento da diversificação dos terópodes no continente sul-americano. O material representa o primeiro registro osteológico de Noasauridae para o Brasil, o segundo representante de noassaurídeo de idade Aptiana, e é também a primeira descrição de um terópode para a região praticamente inexplorada da Bacia Sanfranciscana
The region of the São Francisco craton has a history of sedimentation events dating from the Paleo-Proterozoic and Mesozoic. Sanfranciscana Basin sedimentation represents the last geodynamic cycle that affected the large Sanfranciscana Intracratonic Basin. Despite the Cretaceous of Brazil being the best represented geological period in terms of vertebrate paleofauna, this kind of record in Sanfranciscana Basin was until recently restricted to coelacanths and fragments of fish from Areado Group. Through fieldwork performed by the Laboratory of Paleontology at the Museu de Zoologia - USP, new fossils of continental tetrapods were found in Formation Quiricó sediments, on the vicinity of the city of Coração de Jesus (MG). Among these materials, we highlight epipodials, mesopodials and metapodials elements attributable to a theropod. This study aimed to describe and compare the osteology of this new specimen, numbered as MZSP-PV 830, with other known abelisauroids and also allocate it phylogenetically among the other theropod taxa irradiation of Ceratosauria clade. In this sense, the analysis of the data matrix allocated the new taxon in the Noasauridae clade, based on the following characteristics: the tabular shape of the lateral malleolus of the tibia; and the narrow shape of the metatarsal II shaft. The present description represents a significant paleontological relevance for understanding the diversification of theropods in South America. This material represents the first osteological record of Noasauridae to Brazil, the second noasaurid representative from the Aptian age, as well is the first description of a theropod from the virtually unexplored region of the Sanfranciscana basin
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Ferreira, Rafael Delcourt de Seixas. "Evolução morfológica de Ceratosauria e Tyrannosauroidea (Dinosauria: Theropoda)." Universidade de São Paulo, 2016. http://www.teses.usp.br/teses/disponiveis/38/38131/tde-09122016-115432/.

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Theropoda foram os dinossauros mais bem sucedidos durante e Era Mesozoica e mantiveram-se bem diversos até os tempos presentes. Dentre eles, Tyrannosauroidea e Ceratosauria foram amplamente distribuídos entre os hemisférios norte e sul durante os Períodos Jurássico e Cretáceo, respectivamente. Ambos foram predadores de topo, com formas basais possuindo crânios pequenos, pescoços compridos, pernas e caudas alongadas; enquanto as formas mais derivadas eram caracterizadas com crânios maiores, pescoços curtos e membros anteriores e caudas reduzidas. Apesar de Tyrannosauroidea não ser próximo filogeneticamente de Ceratosauria, esse padrão evolutivo é encontrado em ambas as linhagens. Na presente tese a evolução de ambos os clados é testada utilizando morfometria geométrica nos elementos cranianos, além de medidas lineares e descrições qualitativas. Foram visitadas no total 17 coleções paleontológicas distribuídas no Brasil, Argentina, EUA, Mongólia e China a fim de coletar dados sobre terópodes. Os espécimes examinados foram fotografados, medidos e descritos. Os dados morfométricos foram tratados utilizando os softawares da série TPS e MorphoJ resultando em 54 landmarks nos caracteres cranianos. A fim de avaliar a relação entre os crânios e os landmarks, foram excluídos alguns landmarks do conjunto de dados original. Todos os conjuntos de dados resultantes mostraram poucas variações no morfoespaço, independente do número de landmarks. Foi concluído que a parte anterior do crânio desenvolve-se independentemente da região posterior, após a fenestra anterorbital, em ambos os clados. Ceratosauria é o clado com maior disparidade entre os terópodes carnívoros, especialmente Carnotaurus sastrei. Tyrannosauroidea apresentaram crânios mais conservativos. A disparidade morfológica está relacionada à antiguidade das linhagens e ampla distribuição geográfica. A análise de regressão linear demonstrou que Tyrannosaurus rex pode apresentar grande amplitude fenotípica durante o desenvolvimento ontogenético, e sugere que Nanotyrannus e Raptorex sejam sinônimos juniores de Tyrannosaurus e Tarbosaurus baatar respectivamente. O desenvolvimento ontogenético de Ceratosauria é difícil de avaliar, no entanto Limusaurus inextricabilis apresenta diversas mudanças cranianas durante a fase de crescimento, sugerindo perda de todos os dentes, migração posterior da fenestra pró-maxilar, aumento da órbita e rostro. Ceratosauria e Tyrannosauroidea aumentam de tamanho durante o desenvolvimento filogenético, mas apesar da estrutura corporal ser semelhante, não há convergência morfológica e funcional entre os dois clados. A única convergência entre Tyrannosauroidea e Ceratosauria parece ser ecológica. Tyrannosauridae gráceis como Gorgosaurus libratus, Alioramus altai e juvenis de Tyrannosaurus poderiam ter funções ecológicas semelhantes à Abelisauridae. Por outro lado, Tyrannosauridae mais robustos como Tyrannosaurus, Tarbosaurus e Daspletosaurus torosus poderiam xi ter um nicho mais amplo. Acerca da distribuição e funções ecológicas de Abelisauridae, esse clado poderia ter empurrado Carcharodontosauridae para a extinção, uma vez que ambos apresentam similares convergências craniodentárias e mecânicas.
Theropods were the most successful dinosaurs during the Mesozoic Era, being still well diversied until recent times through the clade Aves. Among them, Tyrannosauroidea and Ceratosauria had a wide distribution, being commonly found on the Northern and Southern Hemispheres, respectively, in the Jurassic and Cretaceous Periods. Both were top predators, known in its basal form, to have small heads, long necks, long forelimbs and tails, whereas the most derived forms were characterized by a large head, short neck, short forelimb and tail. Although Tyrannosauroidea is not phylogenetically close to Ceratosauria, a similar developmental pattern is found in both lineages. In the present thesis, I assessed the evolutionary pattern of morphological shape in both clades using landmark-based geometric morphometric techniques on their skulls, along with some linear measurement and qualitative descriptions. Seventeen paleontological collections distributed in Brazil, Argentina, USA, Mongolia, and China were visited to collect data on these theropods. Specimens examined were photographed, measured and described. The morphometric data were treated using the software TPS series and MorphoJ resulting in 54 landmarks on skull characters. To assess different relationships among the landmarks and the skulls I excluded some landmarks in the original dataset. All datasets show few variations in morphospace, independent of the number of landmarks. Here, I conclude that the anterior part of the skull developed independently from the posterior part of the braincase (posterior to the antorbital fenestra) in both groups independently. Ceratosauria is the clade with more disparity among carnivorous theropods, especially when considering Carnotaurus sastrei. Tyrannosauroidea shows signs of having more conservative skulls. The development of morphological disparity is related to old lineages with long geographical distributions. The linear regression showed that Tyrannosaurus rex could have been more phenotypically spread during ontogenetic development, suggesting that Nanotyrannus and Raptorex are junior synonyms of Tyrannosaurus and Tarbosaurus baatar, respectively. Ontogenetic traits in Ceratosauria are difficult to assess, but Limusaurus inextricabilis showed high rates of change on its skull throughout its development, suggesting that they might loose all teeth, and have xii posterior migration of the promaxillary fenestra, increasing the orbit and snout during the ontogenetic development. Ceratosauria and Tyrannosauroidea show a tendency to larger and heaver forms from basal to more derived forms in their phyogeny. Although main body plan is similar in both group, there are neither morphological nor functional convergences between Ceratosauria and Tyrannosauroidea. The only observed convergence between Tyrannosauridae and Ceratosauria appears to be ecological. Gracile Tyrannosauridae such as Gorgosaurus libratus, Alioramus altai and juvenile Tyrannosaurus may have had similar ecological roles with Abelisauridae. On the other hand, Tyrannosauridae such as Tyrannosaurus, Tarbosaurus and Daspletosaurus torosus may have had a broader niche. Concerning distribution and ecological functions of Abelisauridae, this clade may have pushed another southern clade, the Carcharodontosauridae, to extinction since both clades had similar craniodental and mechanical skull morphologies.
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Bates, Karl T. "Body Size and Locomotor Biomechanics in Non-Avian Theropod Dinosaurs." Thesis, University of Manchester, 2010. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.518454.

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Kambic, Robert Emmett. "Multivariate analysis of avian and non-avian theropod pedal phalanges." Thesis, Montana State University, 2008. http://etd.lib.montana.edu/etd/2008/kambic/KambicR0508.pdf.

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The relationship between morphology and behavior in theropods was examined using multivariate analyses performed on a set of linear measurements of pedal non-ungual phalanges in a sample of 132 extant and 13 extinct taxa. Principal component analysis reveals that modern birds with terrestrial lifestyles tend to cluster away from those with raptorial and non-predatory grasping lifestyles although the division is not clean. Most non-avian dinosaurs tend to cluster with terrestrial species although Deinonychus and some Tyrannosaurids cluster with raptorial taxa. Terrestrial taxa tend to have phalanges that are comparatively shorter distally, are relatively wide, and have shallow grooving of the distal trochleae, while grasping taxa show opposing trends. Predatory graspers have proportionately wider phalanges than non-predatory graspers. Discriminant function analysis performs well in distinguishing among species with specialized behavior while taxa with less stereotyped behaviors are harder to classify. Predatory graspers are easily separated from non-predatory graspers. Troodon and Elmisaurus are grouped with terrestrial/cursorial taxa. Tyrannosaurus, Allosaurus, Daspletosaurus, and Albertosaurus are identified as terrestrial/cursorial with some possible predatory grasping ability. Deinonychus and Bambiraptor appear to have more grasping ability than Troodon.
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Sales, Marcos André Fontenele. "Contribuições à paleontologia de Terópodes não-avianos do Mesocretáceo do Nordeste do Brasil." reponame:Biblioteca Digital de Teses e Dissertações da UFRGS, 2017. http://hdl.handle.net/10183/181014.

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A Região Nordeste do Brasil apresenta um importante registro fóssil de dinossauros mesocretáceos, especialmente do clado Theropoda, popularmente referidos como terópodes. A quase totalidade dos espécimes é oriunda de dois depósitos sedimentares, a Formação Romualdo, Albiano da Bacia do Araripe, e a Formação Alcântara, Cenomaniano da Bacia de São Luís-Grajaú. A presente tese apresenta o estado da arte referente aos terópodes não-avianos das formações supracitadas, contextualizando-os do ponto de vista paleogeográfico e paleoecológico. A revisão dos principais achados é acompanhada por resultados inéditos obtidos pelo pós-graduando ao longo de seu Doutorado, dos quais os principais estão sumarizados a seguir: (1) a obtenção de evidência estatística para a associação positiva entre espinossaurídeos e paleoambientes costeiros; (2) o primeiro registro inequívoco do táxon Abelisauridae na Formação Alcântara; (3) o reconhecimento de uma distinção faunística entre a diversidade dinossauriana das formações Romualdo e Alcântara; (4) a identificação das coroas dentárias maxilares do espinossaurídeo Irritator challengeri; (5) a obtenção de uma nova hipótese para as relações filogenéticas entre os espinossaurídeos brasileiros e os demais táxons. Estes resultados, acompanhados de outros, possuem implicações taxonômicas, evolutivas, paleobiogeográficas e paleoecológicas. Por exemplo, a maior abundância de somatofósseis de espinossaurídeos nas formações mencionadas provavelmente está relacionada à associação positiva entre estes terópodes e os paleoambientes costeiros. Por outro lado, a identificação das coroas dentárias maxilares de I. challengeri, em conjunto com os resultados cladísticos, permitiram a reapreciação de detalhes relativos à evolução das narinas externas de espinossaurídeos, bem como a refutação da hipótese de que o holótipo desta espécie é parte do mesmo espécime do holótipo de Angaturama limai. Ademais, a análise cladística realizada sugere que o clado Baryonychinae não é monofilético. Por fim, o reconhecimento da presença de abelissaurídeos na Formação Alcântara está de acordo com a abundância deste táxon em depósitos sedimentares gondwânicos durante o mesocretáceo, reforçando as semelhanças existentes com os depósitos mesocretáceos africanos, ao mesmo tempo em que salienta as diferenças entre esta unidade e a Formação Romualdo quanto à diversidade de dinossauros. Isto sugere que a fauna dinossauriana da América do Sul era mais heterogênea do que previamente assumido por outros autores, o que deve ser levado em consideração quando de análises futuras de similaridade de faunas. No entanto, a despeito do que já se conhecia e dos novos dados aportados por esta tese, muitas outras questões ainda permanecem por serem investigadas mais a fundo. Desta forma, esta contribuição se constitui em um passo adicional – porém nunca definitivo – no propósito de expandir o conhecimento sobre os terópodes não-avianos do mesocretáceo do Nordeste do Brasil.
The northeastern Brazil has yielded an important fossil record of Mid- Cretaceous dinosaurs, especially the clade Theropoda. Almost all specimens come from two fossiliferous deposits, the Romualdo Formation, Albian of the Araripe Basin, and the Alcântara Formation, Cenomanian of the São Luis-Grajaú Basin. This thesis presents the state of the art relative to the nonavian theropods from the aforementioned formations, contextualizing them paleobiogeographically and paleoecologically. The review of the findings is followed by the inedit results gathered by the PhD candidate during his Doctorate, among which the main ones are summarized below: (1) the obtention of statistical evidence of the positive association between spinosaurids and coastal paleoenvironments; (2) the first unequivocal record of Abelisauridae from the Alcântara Formation; (3) the recognition of a faunal distinction between the dinosaur diversity of the Alcântara and Romualdo formations; (4) the identification of the maxillary tooth crowns of the spinosaurid Irritator challengeri; (5) the obtention of a new hypothesis of the phylogenetic relationships between the Brazilian spinosaurids and the other taxa. These results, along with others, have taxonomic, evolutionary, paleobiogeographic and paleoecological implications. For example, the greater abundance of spinosaurid body fossils in the those formations are likely related to the positive association between these theropods and coastal paleoenvironments. On the other hand, the identification of the maxillary tooth crowns of I. challengeri, along with the cladistic results, enabled the reanalysis of some details of the evolution of the external nares of spinosaurids, as well as the refutation of the hypothesis of the holotypes of this species and Angaturama limai belonging to the same specimen. Furthermore, the cladistic analysis suggests that Baryonychinae is not monophyletic. Finally, the record of abelisaurids from the Alcântara Formation is in agreement with the abundance of this taxon in Gondwanan deposits during the mid-Cretaceous, which strengthens the similarities with African mid-Cretaceous deposits, whereas it emphasizes the differences in dinosaur diversity between this unity and the Romualdo Formation. This suggests that the dinosaur fauna of South America was more heterogeneous than previously assumed by other authors, which must be taken into account by future analyses of similarities. Despite what was previously known and the new findings provided by this thesis, many other issues are still to be further investigated. Thus, this contribution is actually an additional step – although not the definite one – towards the expansion of the knowledge on mid-Cretaceous nonavian theropods from northeastern Brazil.
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Griffin, Christopher T. "The evolution of intraspecific variation, growth, and body size in early theropod dinosaurs." Thesis, Virginia Tech, 2016. http://hdl.handle.net/10919/73682.

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Understanding the changes undergone during the life of an organism is often crucial to properly interpreting the evolutionary history of a group. For extinct organisms, this process can only be directly studied through growth series of fossils representing individuals at different stages of maturity. The growth patterns of the earliest dinosaurs (230–190 million years ago), in particular the morphological changes undergone during the life history of an individual (i.e., ontogeny) is poorly understood. To tackle this problem, I studied the changes undergone during growth of two early theropod dinosaurs, Coelophysis bauri and Megapnosaurus rhodesiensis. To reconstruct the growth of these dinosaurs I used ontogenetic sequence analysis (OSA). I found that, unlike living birds, early dinosaurs possessed an extremely high amount of intraspecific variation in growth. This variation had been previously interpreted as sexual difference; however, I found no evidence of this. Because this variation is widespread among early dinosaurs and their relatives, I hypothesize that this is the ancestral condition of dinosaurian growth, and that this was lost along the evolution to birds. These ontogenetic events are conserved through evolution, and I used this to assess the maturity of large Triassic theropods: I suggest that all known large-bodied Triassic theropods were still growing rapidly at death, and that the maximum body size of Triassic theropods was higher than previously supposed. Theropods were large before the end Triassic mass extinction, unlike what has been previously hypothesized.
Master of Science
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Books on the topic "Theropods"

1

World Book, Inc. Fierce dinosaurs: The large Theropods. Chicago: World Book, 2013.

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Fierce dinosaurs: The large Theropods. Chicago: World Book, 2013.

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World Book, Inc. Birdlike dinosaurs: Small Theropods and prehistoric birds. Chicago: World Book, 2013.

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Birdlike dinosaurs: Small Theropods and prehistoric birds. Chicago: World Book, 2013.

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Naish, Darren William. Studies on Wealden Group theropods: An investigation into the historical taxonomy and phylogenetic affinities of new and previously neglected specimens. Portsmouth: University of Portsmouth, School of Earth, Environmental and Physical Sciences, 1999.

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Lambe, Lawrence M. The Cretaceous Theropodus dinosaur Gorgosaurus. Ottawa: Govt. Print. Bureau, 1997.

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Madsen, James H. Ceratosaurus (Dinosauria, Theropoda): A revised osteology. [Salt Lake City]: Utah Geological Survey, 2000.

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Association, Palaeontological, ed. The interrelationships and evolution of basal theropod dinosaurs. London: Palaeontological Assoc., 2003.

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Hutt, Stephen. Neovenator salerii: A new theropod dinosaur from the wealden of the Isle of Wight : its status and significance for theropod evolution. Portsmouth: University of Portsmouth, School of Earth, Environmental and Physical Sciences, 1999.

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Livezey, Bradley C. Higher-order phylogeny of modern birds (Theropoda, Aves:Neornithes) based on comparative anatomy. Pittsburgh, Pa: Carnegie Museum of Natural History, 2006.

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Book chapters on the topic "Theropods"

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Prothero, Donald R. "Theropods." In Vertebrate Evolution, 265–88. Boca Raton: CRC Press, 2022. http://dx.doi.org/10.1201/9781003128205-18.

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Agnolín, Federico L., and Fernando E. Novas. "Uncertain Averaptoran Theropods." In SpringerBriefs in Earth System Sciences, 37–47. Dordrecht: Springer Netherlands, 2013. http://dx.doi.org/10.1007/978-94-007-5637-3_4.

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Xu, Xing. "Filamentous Integuments in Nonavialan Theropods and Their Kin: Advances and Future Perspectives for Understanding the Evolution of Feathers." In The Evolution of Feathers, 67–78. Cham: Springer International Publishing, 2020. http://dx.doi.org/10.1007/978-3-030-27223-4_5.

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Makovicky, Peter J., and Lindsay E. Zanno. "Theropod Diversity and the Refinement of Avian Characteristics." In Living Dinosaurs, 9–29. Chichester, UK: John Wiley & Sons, Ltd, 2011. http://dx.doi.org/10.1002/9781119990475.ch1.

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Oettl-Rieser, Andrea B., and Marion Zahner. "FRICK: Late Triassic Basal Sauropodomorph and Theropod Dinosaurs at the Sauriermuseum Frick, Switzerland." In Paleontological Collections of Germany, Austria and Switzerland, 241–48. Cham: Springer International Publishing, 2018. http://dx.doi.org/10.1007/978-3-319-77401-5_22.

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Prasad, Guntupalli V. R., and Varun Parmar. "First Ornithischian and Theropod Dinosaur Teeth from the Middle Jurassic Kota Formation of India: Paleobiogeographic Relationships." In Biological Consequences of Plate Tectonics, 1–30. Cham: Springer International Publishing, 2020. http://dx.doi.org/10.1007/978-3-030-49753-8_1.

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Novas, Fernando E., Sankar Chatterjee, Dhiraj K. Rudra, and P. M. Datta. "Rahiolisaurus gujaratensis, n. gen. n. sp., A New Abelisaurid Theropod from the Late Cretaceous of India." In New Aspects of Mesozoic Biodiversity, 45–62. Berlin, Heidelberg: Springer Berlin Heidelberg, 2010. http://dx.doi.org/10.1007/978-3-642-10311-7_3.

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Chatterjee, S. "A New Theropod Dinosaur from India with Remarks on the Gondwana-Laurasia Connection in the Late Triassic." In Gondwana Six: Stratigraphy, Sedimentology, and Paleontology, 183–89. Washington, D. C.: American Geophysical Union, 2013. http://dx.doi.org/10.1029/gm041p0183.

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Marugán-Lobón, Jesús, and Ángela D. Buscalioni. "Geometric morphometrics in macroevolution: morphological diversity of the skull in modern avian forms in contrast to some theropod dinosaurs." In Morphometrics, 157–73. Berlin, Heidelberg: Springer Berlin Heidelberg, 2004. http://dx.doi.org/10.1007/978-3-662-08865-4_12.

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Queiroz, Kevin de, Philip D. Cantino, and Jacques A. Gauthier. "Theropoda O. C. Marsh 1881 [D. Naish, A. Cau, T. R. Holtz, Jr., M. Fabbri, and J. A. Gauthier], converted clade name." In Phylonyms, 1235–46. Boca Raton : CRC Press, [2019]: CRC Press, 2020. http://dx.doi.org/10.1201/9780429446276-282.

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Conference papers on the topic "Theropods"

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Rhoda, Daniel, Spencer Hellert, and P. David Polly. "SHIFTING PATTERNS OF FUNCTIONAL INTEGRATION DURING THE EVOLUTION OF FLIGHT IN THEROPODS." In GSA Annual Meeting in Indianapolis, Indiana, USA - 2018. Geological Society of America, 2018. http://dx.doi.org/10.1130/abs/2018am-323639.

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Spendlove, Karrah, Andrew R. C. Milner, Jerald D. Harris, and Randall Irmis. "GRALLATOR TRACKS AS A WINDOW TO GROWTH STRATEGIES OF SMALL-BODIED EARLY THEROPODS." In Joint 118th Annual Cordilleran/72nd Annual Rocky Mountain Section Meeting - 2022. Geological Society of America, 2022. http://dx.doi.org/10.1130/abs/2022cd-374121.

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Breithaupt, Brent H., Anne Schulp, Neffra A. Matthews, Martin Lockley, Richard T. McCrea, Lisa G. Buckley, Gerard Gierlinski, et al. "NEST SCRAPE BEHAVIOR IN LARGE THEROPODS: FROM COLORADO OUTCROP TO DUTCH SHOWCASE: VISITORS EXPERIENCE ANCIENT DINOSAUR DISPLAY BEHAVIOR FROM FOSSIL EVIDENCE." In GSA Annual Meeting in Seattle, Washington, USA - 2017. Geological Society of America, 2017. http://dx.doi.org/10.1130/abs/2017am-306527.

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Breithaupt, Brent, Neffra A. Matthews, ReBecca Hunt-Foster, H. Gregory McDonald, and Martin Lockley. "CHASING THEROPODS ACROSS AN EARLY JURASSIC SAND SEA: “PALEOCAMP” OPPORTUNITIES FOR YOUNG CITIZEN SCIENTISTS TO HELP DISCOVER, DOCUMENT, INTERPRET, MANAGE, AND PRESERVE AN ANCIENT TRACKSITE IN THE NAVAJO SANDSTONE OF SOUTHEASTERN UTAH." In GSA Connects 2021 in Portland, Oregon. Geological Society of America, 2021. http://dx.doi.org/10.1130/abs/2021am-368233.

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Frederico, Charles Austin, and Matthew McLain. "NEW CAENAGNATHID (THEROPODA: OVIRAPTOROSAURIA) MATERIAL FROM THE LANCE FORMATION (MAASTRICHTIAN) OF WYOMING." In GSA Connects 2021 in Portland, Oregon. Geological Society of America, 2021. http://dx.doi.org/10.1130/abs/2021am-370156.

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Buntin, Rogers, Neffra A. Matthews, Brent H. Breithaupt, Martin Lockley, and Nora Noffke. "THEROPOD COURTSHIP BEHAVIOR PRESERVED BY MICROBIAL MATS, CRETACEOUS DAKOTA SANDSTONE, COLORADO, USA." In GSA Connects 2022 meeting in Denver, Colorado. Geological Society of America, 2022. http://dx.doi.org/10.1130/abs/2022am-382140.

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Bykowski, Richard, Katlin Schroeder, and P. David Polly. "NON-AVIAN THEROPOD SKULL MORPHOLOGY TENTATIVELY SUPPORTS ONTOGENETIC NICHE SHIFT AMONG NORTH AMERICAN TYRANNOSAURIDS." In GSA Connects 2021 in Portland, Oregon. Geological Society of America, 2021. http://dx.doi.org/10.1130/abs/2021am-368719.

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Heckert, Andrew B., Justin A. Spielmann, Spencer G. Lucas, Richard Altenberg, and Daniel M. Russell. "An Upper Jurassic theropod dinosaur from the Section 19 mine, Morrison Formation, Grants Uranium district." In 54th Annual Fall Field Conference. New Mexico Geological Society, 2003. http://dx.doi.org/10.56577/ffc-54.309.

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Dalman, Sebastian G., Amanda K. Cantrell, Thomas L. Suazo, and Spencer G. Lucas. "First Record of a Tyrannosaurid Theropod (dinosauria) From the Lower Campanian Menefee Formation, New Mexico." In 2016 New Mexico Geological Society Annual Spring Meeting. Socorro, NM: New Mexico Geological Society, 2016. http://dx.doi.org/10.56577/sm-2016.396.

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Williamson, Thomas E., and Stephen L. Brusatte. "Small Theropod Teeth from the Late Cretaceous of the San Juan Basin, Northwestern New Mexico." In 2014 New Mexico Geological Society Annual Spring Meeting. Socorro, NM: New Mexico Geological Society, 2014. http://dx.doi.org/10.56577/sm-2014.242.

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