Relevant bibliographies by topics / Tetraploid wheat

Academic literature on the topic 'Tetraploid wheat'

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Published: 10 December 2022
Last updated: 28 January 2023

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Journal articles on the topic "Tetraploid wheat"

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Zhangaziev, A. S., S. I. Nurbekov, G. K. Ziyaeva, and J. S. Tulubaev. "Origin of cultivated species of tetraploid wheat." International Journal of Biology and Chemistry 7, no. 2 (2014): 57–60. http://dx.doi.org/10.26577/2218-7979-2014-7-2-57-60.

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Joppa, L. R. "Chromosome Engineering in Tetraploid Wheat." Crop Science 33, no. 5 (September 1993): 908–13. http://dx.doi.org/10.2135/cropsci1993.0011183x003300050006x.

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Pronozin, A. Yu, A. A. Paulish, E. A. Zavarzin, A. Yu Prikhodko, N. M. Prokhoshin, Yu V. Kruchinina, N. P. Goncharov, E. G. Komyshev, and M. A. Genaev. "Automatic morphology phenotyping of tetra- and hexaploid wheat spike using computer vision methods." Vavilov Journal of Genetics and Breeding 25, no. 1 (March 16, 2021): 71–81. http://dx.doi.org/10.18699/vj21.009.

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Intraspecific classification of cultivated plants is necessary for the conservation of biological diversity, study of their origin and their phylogeny. The modern cultivated wheat species originated from three wild diploid ancestors as a result of several rounds of genome doubling and are represented by di-, tetra- and hexaploid species. The identification of wheat ploidy level is one of the main stages of their taxonomy. Such classification is possible based on visual analysis of the wheat spike traits. The aim of this study is to investigate the morphological characteristics of spikes for hexa- and tetraploid wheat species based on the method of high-performance phenotyping. Phenotyping of the quantitative characteristics of the spike of 17 wheat species (595 plants, 3348 images), including eight tetraploids (Triticum aethiopicum, T. dicoccoides, T. dicoccum, T. durum, T. militinae, T. polonicum, T. timopheevii, and T. turgidum) and nine hexaploids (T. compactum, T. aestivum, i:ANK-23 (near-isogenic line of T. aestivum cv. Novosibirskaya 67), T. antiquorum, T. spelta (including cv. Rother Sommer Kolben), T. petropavlovskyi, T. yunnanense, T. macha, T. sphaerococcum, and T. vavilovii), was performed. Wheat spike morphology was described on the basis of nine quantitative traits including shape, size and awns area of the spike. The traits were obtained as a result of image analysis using the WERecognizer program. A cluster analysis of plants according to the characteristics of the spike shape and comparison of their distributions in tetraploid and hexaploid species showed a higher variability of traits in hexaploid species compared to tetraploid ones. At the same time, the species themselves form two clusters in the visual characteristics of the spike. One type is predominantly hexaploid species (with the exception of one tetraploid, T. dicoccoides). The other group includes tetraploid ones (with the exception of three hexaploid ones, T. compactum, T. antiquorum, T. sphaerococcum, and i:ANK-23). Thus, it has been shown that the morphological characteristics of spikes for hexaploid and tetraploid wheat species, obtained on the basis of computer analysis of images, include differences, which are further used to develop methods for plant classifications by ploidy level and their species in an automatic mode.
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Sarrafi, A., N. Amrani, and G. Alibert. "Haploid regeneration from tetraploid wheat using maize pollen." Genome 37, no. 1 (February 1, 1994): 176–78. http://dx.doi.org/10.1139/g94-023.

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Crosses were made between 21 tetraploid wheat genotypes (6 parents, 15 F1 hybrids) and a single F1 hybrid of maize that was used as the male parent. Plants were grown under controlled greenhouse conditions (daylength, 16 h; temperature, 25 °C days and 15 °C nights). To enhance embryo survival, 2,4-D (10 mg/L) was applied to spikes 24 h after pollination with maize. Embryos were recovered from the tetraploid wheat genotypes at a rate of 2.34–14.14/100 developed ovaries. Sixty-nine haploid plants were obtained from 3 parents and 12 F1, hybrids. Fifty-six of these were successfully doubled. General combining ability was significant for the two traits studied, indicating that additive genetic control is important for the number of developed ovaries and haploid embryo production in tetraploid wheat × maize crosses. In this report, we demonstrate the potential of using maize pollen to produce haploid plants from tetraploid wheat genotypes.Key words: tetraploid wheat, embryo culture, haploid, wheat × maize, combining abilities.
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Joshi, Chandrashekhar P., and Henry T. Nguyen. "Application of the random amplified polymorphic DNA technique for the detection of polymorphism among wild and cultivated tetraploid wheats." Genome 36, no. 3 (June 1, 1993): 602–9. http://dx.doi.org/10.1139/g93-081.

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Development of a high-density genetic linkage map of cultivated wheats using conventional molecular markers has lagged behind the other major food crops such as rice and tomato because of the large genome size and limited levels of genetic polymorphisms. Recently, random amplified polymorphic DNAs (RAPDs) have been suggested to provide an alternative to visualize more polymorphism. For the construction of a genetic linkage map in tetraploid wheats, one can use a strategy of intersubspecific crosses between the most dissimilar wild and cultivated tetraploid wheats that are easy to hybridize and result in fertile progeny. An assessment of the level of RAPDs among different accessions and varieties of wild and cultivated tetraploid wheats is required to fulfill this objective. We present here the data obtained using RAPD analysis of 40 primers in 20 accessions of wild tetraploid emmer wheats (Triticum turgidum L. ssp. dicoccoides) and 10 genotypes of cultivated tetraploid durum wheats (Triticum turgidum L. ssp. durum) selected from geographically diverse locations. We have observed a higher level of polymorphism among different accessions of wild emmer wheat from Israel, Turkey, and Jordan than the group of cultivated American, Turkish, and Syrian durum wheats. These data have been used to generate a dendrogram suggesting the genetic relationships among these genotypes, and the most dissimilar genotypes are identified for future mapping and gene tagging work.Key words: durum wheat, emmer wheat, genetic similarity, molecular markers, RAPD analysis.
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Zhao, Na, Qianli Dong, Brian D. Nadon, Xiaoyang Ding, Xutong Wang, Yuzhu Dong, Bao Liu, Scott A. Jackson, and Chunming Xu. "Evolution of Homeologous Gene Expression in Polyploid Wheat." Genes 11, no. 12 (November 25, 2020): 1401. http://dx.doi.org/10.3390/genes11121401.

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Polyploidization has played a prominent role in the evolutionary history of plants. Two recent and sequential allopolyploidization events have resulted in the formation of wheat species with different ploidies, and which provide a model to study the effects of polyploidization on the evolution of gene expression. In this study, we identified differentially expressed genes (DEGs) between four BBAA tetraploid wheats of three different ploidy backgrounds. DEGs were found to be unevenly distributed among functional categories and duplication modes. We observed more DEGs in the extracted tetraploid wheat (ETW) than in natural tetraploid wheats (TD and TTR13) as compared to a synthetic tetraploid (AT2). Furthermore, DEGs showed higher Ka/Ks ratios than those that did not show expression changes (non-DEGs) between genotypes, indicating DEGs and non-DEGs experienced different selection pressures. For A-B homeolog pairs with DEGs, most of them had only one differentially expressed copy, however, when both copies of a homeolog pair were DEGs, the A and B copies were more likely to be regulated to the same direction. Our results suggest that both cis- and inter-subgenome trans-regulatory changes are important drivers in the evolution of homeologous gene expression in polyploid wheat, with ploidy playing a significant role in the process.
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Allaby, Robin G., and Terence A. Brown. "Reply to the comment by Salamini et al. on "AFLP data and the origins of domesticated crops"." Genome 47, no. 3 (June 1, 2004): 621–22. http://dx.doi.org/10.1139/g04-012.

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We reiterate the key points of a previous paper that showed that neighbor-joining analysis of AFLP datasets can produce erroneous results. The critical question, whether the datasets used to infer the origins of einkorn, barley, and the hulled and hard tetraploid wheats display sufficient linkage to avoid the artifacts that we observed, is not adequately answered by Salamini et al.Key words: AFLPs, crop domestication, einkorn wheat, barley, tetraploid wheat.
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Muramatsu, Mikio. "The vulgare super gene, Q: its universality in durum wheat and its phenotypic effects in tetraploid and hexaploid wheats." Canadian Journal of Genetics and Cytology 28, no. 1 (February 1, 1986): 30–41. http://dx.doi.org/10.1139/g86-006.

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The genotype at the Q locus on chromosome 5A of Triticum durum Desf. (2n = 28, AABB), a species with keeled glumes and tough rachis, was studied by either crossing the species with, or substituting its 5A into, a hexaploid common wheat, T. aestivum (L.)Thell. ssp. vulgare (Vill.) MK. cv. Chinese Spring (2n = 42, AABBDD, QQ genotype). Contrary to the opinion of previous researchers that keeled-glumed wheats always have the spelta gene, q, the durum strains studied had a hypermorphic allele, the vulgare gene, Q. No speltoid plants appeared in the progeny of the crosses, and disomic substitution lines (2n = 2) had squareheaded (= vulgare type) spikes. Also, three doses of the long arm of 5A of durum produced compactoidy. Apparently, Q does not produce round glumes in the genetic backgrounds of most tetraploids except T. carthlicum Nevski. The phenotype conditioned by Q, which is evidently present in all free-threshing tetraploid wheats, is somewhat different at the tetraploid level from that in hexaploids. The presence of Q tends to magnify the differences in the degree of expression of certain minor characters.Key words: Triticum, phylogeny, vulgare gene, pleiotropic gene, dosage effect, interaction.
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Lelley, T., E. Kazman, K. M. Devos, and M. D. Gale. "Use of RFLPs to determine the chromosome composition of tetraploid triticale (A/B)(A/B)RR." Genome 38, no. 2 (April 1, 1995): 250–54. http://dx.doi.org/10.1139/g95-031.

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Tetraploid triticale, (A/B)(A/B)RR (2n = 28), is a botanical novelty, an amphiploid composed of a diploid rye and a 14 chromosome wheat genome made up of chromosomes of the A and B genomes of tetraploid wheat. Restriction fragment length polymorphism (RFLP) markers were used to elucidate the chromosome composition of the mixed wheat genome of 35 different tetraploid triticale lines. Of 128 possible A/B chromosome pair combinations, only 6 were found among these lines, with a prevalence of the 1A, 2A, 3B, 4B, 5B, 6B, and 7B karyotype. In most triticale lines stable wheat genomes made up of only homologous A or B genome chromosome pairs were identified, however, in some lines homoeologous chromosome pairs were found. In this paper we demonstrate that RFLPs can be used successfully as an alternative to C-banding for the identification of the chromosome composition of tetraploid triticale and discuss the possible selective advantage of specific chromosome composition.Key words: tetraploid triticale, mixed wheat genome, RFLR
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Wang, H., and J. M. Clarke. "Relationship of excised-leaf water loss and stomatal frequency in wheat." Canadian Journal of Plant Science 73, no. 1 (January 1, 1993): 93–99. http://dx.doi.org/10.4141/cjps93-012.

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Rate of water loss from excised leaves of wheat (Triticum spp.) is associated with adaptation to dry growing conditions, but the causes of observed genotypic differences are not well understood. This study was conducted to determine the relationship between stomatal characteristics and excised-leaf water status in tetraploid (Triticum turgidum L. var. durum) and hexaploid (Triticum aestivum L.) wheat genotypes. Samples were taken from field and growth-room experiments to measure stomatal frequency (SF) and size, leaf water content at excision (WC0) and 30 min after excision (WC30), rate of water loss (RWL) 30-120 min after excision, epidermal conductance (ge), and relative water content (RWC). SF was not correlated with RWL in the field experiments and was negatively correlated with WC0 and WC30 in tetraploids but not in hexaploids. In the growth-room experiment, SF was positively correlated with ge 50 and 30 min after excision for tetraploid and hexaploid genotypes, respectively. SF was correlated with RWL in tetraploids (r = 0.64*, n = 12) and hexaploids (r = 0.81**, n = 12). However, there were no significant correlations between stomatal characteristics and WC0, WC30 or RWC. These results indicate that SF is perhaps one of several factors influencing genotypic differences in excised-leaf water loss. The inconsistency of this relationship may be due to the influence of other traits affecting RWL. Key words: Leaf water loss, stomata, drought, Triticum aestivum L., T. turgidum L. var. durum
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Dissertations / Theses on the topic "Tetraploid wheat"

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Tavakkol, Afshari Reza. "Variation in seed dormancy of tetraploid wheat." Thesis, National Library of Canada = Bibliothèque nationale du Canada, 1999. http://www.collectionscanada.ca/obj/s4/f2/dsk2/ftp03/NQ37916.pdf.

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Khan, Javed Ahmad. "Salinity effects on 4D recombinant tetraploid wheat genotypes." Thesis, Bangor University, 1996. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.321525.

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Liu, Chao-yin. "Variation and genetic control of prolamins in tetraploid wheats and their association with quality in durum wheat." Title page, contents and summary only, 1994. http://web4.library.adelaide.edu.au/theses/09PH/09phl783.pdf.

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Maddi, Satyanarayana. "DNA-based food authentication techniques : differentiation of tetraploid and hexaploid wheat." Thesis, Glasgow Caledonian University, 2010. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.517961.

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Saini, Jyoti. "Genomic Analysis of Domestication-Related Traits and Stem Rust Resistance in Tetraploid Wheat." Diss., North Dakota State University, 2017. https://hdl.handle.net/10365/27459.

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Modern durum and common wheat cultivars were developed from ancient wheat ancestors by natural and artificial selection of agronomic and domestication traits, which ultimatey decreased their genetic diversity and made them more susceptible to various biotic and abiotic stresses. At present, new sources of resistance need to be introgressed into future wheat cultivars to combat the effect of the disease stem rust caused by the biotrophic fungal pathogen Puccinia graminis f.sp. tritici (Pgt). In this dissertation, I first analyzed the domestication-related traits in a tetraploid recombinant inbred line (RIL) population developed from a cross between the durum wheat line Rusty and the cultivated emmer accession PI 193883 (referred to as the RP883 population). Second, the RP883 population and a double haploid (DH) population (referred to as the LP749 population) derived from a cross between the durum cultivar Lebsock and the Triticum. turgidium ssp. carthlicum accession PI 94749, and nine durum wheat cultivars were screened with Pgt races TMLKC, TTKSK, TRTTF, and TTTTF. Domestication-related trait analysis in the RP883 population showed vernalization (Vrn-A1) and domestication (Q) genes had a pleiotrophic effect on spike length, spikelet per spike, spike compactness, and threshability. Additionally, an interaction and dosage effect of three free-threshing trait governing loci, teneacious glume Tg2A and Tg2B, and q, revealed that mutation in all three loci are required to attain complete free threshability. The stem rust analysis done in the RP883 population showed two Sr gene regions conferring resistance against TMLKC, TTKSK, and TRTTF: one novel gene region on chromosome 2BL (Sr883) and likely a new allele or gene residing in close proximity to the Sr13 gene on 6AL. The second stem rust study using the LP749 population and nine durum wheat cultivars showed that most likely the U.S. durum germplasm carries the four major Sr genes, Sr7a (4AL), Sr8155B1 (6AS), Sr13 (6AL), and likely Sr9e (2BL) against TTKSK, TRTTF, and TTTTF. In conclusion, results obtained from this domestication study provide knowledge about different stages in wheat evolution. Both stem rust studies revealed genetic diversity in the tetraploid wheat gene pool and indicate their utility in future breeding programs.
USDA-Agricultural Research Service
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Oliveira, Hugo Rafael Cardoso. "Archaeogenetics and the spread of agriculture in the Iberian Peninsula and Northwest Africa : a study of genetic variation within tetraploid and diploid wheats." Thesis, University of Cambridge, 2011. http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.609794.

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Rahman, Muhammad Shefatur. "Genetic control of sodium exclusion in tetraploid wheat." Thesis, 2011. http://hdl.handle.net/2440/82381.

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Worldwide, soil salinity is one of the major abiotic stress factors limiting crop production. Durum wheat (Triticum turgidum L. ssp. durum [Desf.]) is a relatively salt sensitive species. Its sensitivity to salt is thought to be due to its poor ability in limiting accumulation of toxic Na⁺ in leaf tissues. The present research was undertaken to investigate the genetic control of sodium exclusion in a tetraploid wheat population of recombinant inbred lines developed from a durum wheat cultivar Atil and a cultivated emmer wheat (Triticum turgidum L. ssp. dicoccum [Schrank].Thell.) accession PI94628. In order to estimate the nature and extent of variation for sodium exclusion within the progeny, two experiments were carried out using a supported hydroponic system. In both cases, Na⁺ and K⁺ concentration were determined from the fourth leaf of plants which had imposed a 100 mM NaCl stress for 10 days. In the first experiment, screening of replicated parental lines and a subset of inbred lines (24 RILs) indicated the existence of significant genetic variation for sodium exclusion within the population. It was also found that the spatial variation within the experimental equipment contributed only 8-9% of the total observed phenotypic variation. In the second experiment, screening of the entire population indicated transgressive segregation for both of the Na⁺ and K⁺ accumulation traits, with the durum wheat line Atil found to be the better sodium excluder. A significant negative correlation (r = - 0.7) was found between leaf Na⁺ and K⁺ concentration, however, neither of these traits was found to be correlated with the shoot biomass of 30-day-old seedlings (21 days under salt stress). To construct a genetic linkage map, 1057 markers (916 DArT and 141 SSRs) were used. Of these, 467 markers were eliminated from the linkage analysis due the segregation distortion. The remaining 495 DArTs and 95 SSR markers were used in map construction. They provided reasonable genome coverage (2136.5 cM), with a marker density of 3.6 cM/marker. The markers were distributed on 34 linkage groups representing parts of 14 chromosomes, with gaps of greater than 15 cM still remaining in 12 of the chromosomes. The majority of these markers showed conserved locations and orientation when compared to those described in previous genetic linkage maps of tetraploid and hexaploid wheat. Marker polymorphisms in two regions on chromosome 5A and 5B were significantly associated phenotypic variation for both Na⁺ and K⁺ accumulation in the leaf tissues. In both of these QTL regions, the durum wheat parent was contributed the alleles for lower Na⁺ concentration and higher K⁺ concentration. The two QTLs explained 28% and 19% of total phenotypic variation for Na⁺ and K⁺ accumulation, respectively. Both QTLs were mapped in the centromeric regions of the chromosomes. No QTLs for these traits have previously been reported in these regions. A highly significant QTL for shoot biomass was also mapped on chromosome 5A; explaining 16% of the phenotypic variation but this was over 32.4 cM from the leaf Na⁺ or K⁺ concentration QTLs. Furthermore, the QTLs were not found to be associated with traits related to vigour and/or vernalisation requirement. The molecular markers in the QTL regions detected here could serve as starting points for further characterisation of these genomic regions to elucidate the physiological and molecular bases of these QTLs.
Thesis (M.Ag.Sc.) -- University of Adelaide, School of Agriculture, Food and Wine, 2011
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Sykes, Ellen Elizabeth. "Inheritance of tan spot resistance in hexaploid, tetraploid, and diploid wheat." 1989. http://hdl.handle.net/1993/16973.

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Ali, Mohamed Badry Mohamed. "Elucidating and Mapping Heat Tolerance in Wild Tetraploid Wheat (Triticum turgidum L.)." Thesis, 2010. http://hdl.handle.net/1969.1/ETD-TAMU-2010-12-8888.

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Identifying reliable screening tools and characterizing tolerant germplasm sources is essential for developing wheat (Triticum aestivum L.) varieties suited for the hot areas of the world. Our objective was to evaluate heat tolerance of promising wild tetraploid wheat (Triticum turgidum L.) accessions that could be used as sources of heat tolerance in common- and durum-wheat (Triticum durum) breeding programs. We screened 109 wild tetraploid wheat accessions collected by the International Center for Agriculture Research in the Dry Areas (ICARDA) from the hottest wheat growing areas in Africa and Asia, as well as, two common wheat checks for their response to heat stress by measuring damage to the thylakoid membranes, flag leaf temperature depression (FLTD), and spike temperature depression (STD) during exposure to heat stress for 16 beginning at anthesis. Measurements were taken on the day of anthesis then 4, 8, 12, and 16 days post anthesis (DPA) under controlled optimum and heat-stress conditions. Individual kernel weight (IKW) and heat susceptibility index (HSI) measurements were also obtained. Prolonged exposure to heat stress was associated with increased damage to thylakoid membranes, as indicated by the high ratio of constant fluorescence (O) to peak variable fluorescence (P). A positive and significant correlation was found between O/P ratio and both FLTD and STD under heat-stress conditions. A negative and significant correlation was found between FLTD and HSI and between STD and HSI based on the second and third measurements (4 and 8 DPA). Correlations obtained after the third measurement were not significant because heat-stress accelerated maturity and senescence. For a pedigree-based mapping strategy a family approach was then developed by crossing and back-crossing heat-tolerant and heat-susceptible germplasm. A set of 800 lines resulting from the pedigree-based family approach was phenotyped using FLTD, chlorophyll content and yield and its components under heat stress. Genotyping of these lines was accomplished using simple sequence repeat (SSRs) markers. Some QTLs associated with heat stress tolerance were identified. This study identified potential heat-tolerant wild tetraploid wheat germplasm and QTL conditioning heat tolerance that can be incorporated into wheat breeding programs to improve cultivated common and durum wheat.
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Liu, C. Y. "Variation and genetic control of prolamins in tetraploid wheats and their association with quality in durum wheat / by Chao-yin Liu." 1994. http://hdl.handle.net/2440/21588.

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Bibliography: leaves 180-198.
viii, 217 leaves : ill. (some col.) ; 30 cm.
Title page, contents and abstract only. The complete thesis in print form is available from the University Library.
Thesis (Ph.D.)--University of Adelaide, Dept. of Plant Science, 1994
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Books on the topic "Tetraploid wheat"

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Kosina, Romuald. Tetraploids of the genus Triticum in the light of caryopsis structure. Wrocław: Wydawnictwo Uniwersytetu Wrocławskiego, 1995.

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Book chapters on the topic "Tetraploid wheat"

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Joppa, L. R. "Aneuploid Analysis in Tetraploid Wheat." In Agronomy Monographs, 255–67. Madison, WI, USA: American Society of Agronomy, Crop Science Society of America, Soil Science Society of America, 2015. http://dx.doi.org/10.2134/agronmonogr13.2ed.c11.

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Maccaferri, Marco, Martina Bruschi, and Roberto Tuberosa. "Sequence-Based Marker Assisted Selection in Wheat." In Wheat Improvement, 513–38. Cham: Springer International Publishing, 2022. http://dx.doi.org/10.1007/978-3-030-90673-3_28.

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AbstractWheat improvement has traditionally been conducted by relying on artificial crossing of suitable parental lines followed by selection of the best genetic combinations. At the same time wheat genetic resources have been characterized and exploited with the aim of continuously improving target traits. Over this solid framework, innovations from emerging research disciplines have been progressively added over time: cytogenetics, quantitative genetics, chromosome engineering, mutagenesis, molecular biology and, most recently, comparative, structural, and functional genomics with all the related -omics platforms. Nowadays, the integration of these disciplines coupled with their spectacular technical advances made possible by the sequencing of the entire wheat genome, has ushered us in a new breeding paradigm on how to best leverage the functional variability of genetic stocks and germplasm collections. Molecular techniques first impacted wheat genetics and breeding in the 1980s with the development of restriction fragment length polymorphism (RFLP)-based approaches. Since then, steady progress in sequence-based, marker-assisted selection now allows for an unprecedently accurate ‘breeding by design’ of wheat, progressing further up to the pangenome-based level. This chapter provides an overview of the technologies of the ‘circular genomics era’ which allow breeders to better characterize and more effectively leverage the huge and largely untapped natural variability present in the Triticeae gene pool, particularly at the tetraploid level, and its closest diploid and polyploid ancestors and relatives.
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Al Hakimi, A., P. Monneveaux, and M. M. Nachit. "Direct and indirect selection for drought tolerance in alien tetraploid wheat x durum wheat crosses." In Developments in Plant Breeding, 353–60. Dordrecht: Springer Netherlands, 1997. http://dx.doi.org/10.1007/978-94-011-4896-2_49.

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Prat, Noémie, Maria Buerstmayr, Barbara Steiner, and Hermann Buerstmayr. "Meta-analysis of Resistance to Fusarium Head Blight in Tetraploid Wheat: Implications for Durum Wheat Breeding." In Advances in Wheat Genetics: From Genome to Field, 323–29. Tokyo: Springer Japan, 2015. http://dx.doi.org/10.1007/978-4-431-55675-6_37.

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MUKERJI, K. G. "Mycorrhizal Colonization in Roots of Tetraploid and Hexaploid Wheat Species Suvercha." In Plant Roots and their Environment, 498–505. Elsevier, 1991. http://dx.doi.org/10.1016/b978-0-444-89104-4.50069-4.

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"americanum) [29]. Among wheat, tetraploid durum wheat contained higher FL contents than the U.S. hard winter NSTL shows the highest NL:PoL ratio. wheats. Larsen et al. [66] reported New Zealand wheat flour Among all grains, wheat is the richest in GL, followed FL content ranges of 67-85 mg/10 g (db) for the 1984 crop by triticale, rye, and barley. Millet lipids from P. ameri-and 93-108 mg/10 g (db) for the 1985 wheat crop (Table 4). canum seed [29], corn, and sorghum lipids contain the Ten Greek bread wheat flours [67] contained lipid ranges lowest GL content. However, other researchers [32] report-similar to those in U.S. Kansas flours reported by Chung et ed that GL contents ranged from 6 to 14% for millet lipids al. [61]. Australian scientists [68,69] also investigated their that were extracted by hot water—saturated butanol and wheat FL. Compared with the means of U.S. wheat and acid hydrolysis. flour FL [61], Australian wheats contained substantially In general, PL also are more abundant in wheat, triti-less FL and NL but higher PL. Australian flours contained cale, rye lipids and slightly lower in barley, oat groats, similar FL and NL but still higher PoL content (Table 4). sorghum, and rice. Although corn NSTL were found to have higher PL contents than GL contents, they were very low in PL compared to other grains. Millet NSTL from P. C. Fatty Acid Composition of Grain Lipids americanum seed [29] contains the lowest PL content of All cereal grain lipids are rich in unsaturated fatty acids all the grains. (FA) (Table 5). Palmitic acid (16:0) is a major saturated Wheat flour FL, a minor component, have been report-FA, and linoleic acid (18:2) is a major unsaturated FA for ed to have a significant effect on bread-making. When the all cereals except for brown rice. In brown rice, oleic acid defatted flours were reconstituted with the extracted lipids (18:1) is a major unsaturated FA. The presence of palmi-to their original levels, the PoL fraction of FL but not the toleic acid (16:1) and eicosenoic acid (20:1) is reported NL completely restored loaf volume and crumb grain quite often but usually at levels below 1% of total FA com-[59,60]. Among wheat flour lipids, GL are the best bread position. loaf volume improvers [19-21]. Fatty acid compositions are generally similar for barley, In 1982, Chung et al. [61] reported a range of 177-230 rye, triticale, and wheat lipids. Rye lipids are somewhat mg/10 g (db) for wheat FL contents of 21 HRW wheats higher in linolneic acid (18:3) than those of other cereals. (Table 4). Flours showed 83-109 mg FL, 67-84 mg NL, Oat lipid FA composition is similar to that of brown rice, and 11-27 mg PoL with NL:PoL ratios of 2.5-6.9. Ohm because oats and brown rice are rich in oleic acid. Millet and Chung [62] also investigated the FL contents of flours lipids are generally higher in stearic acid (18:0) than all from 12 commercial hard winter wheat cultivars grown at other cereal lipids. six locations and reported the cultivar mean ranges of There are wide ranges in FA compositions of corn oils 90-109 mg/10 g (db) for total flour FL, 72-85 mg for NL, (Table 6). Jellum [82] reported a range of 14-64% oleic 11-16 mg for GL, 1.7-3.1 mg for monogalactosyldiglyc-acid and 19-71% linoleic acid for the world collection of erides (MGDG), 5.3-6.5 mg for digalactosyldiglycerides 788 varieties of corn (Table 6). The wide ranges in FA com-(DGDG), and 5-7 mg for PL (Table 4). The ratios of NL to position were due to more lines having been examined in PoL were in a much narrower range than those of earlier corn than in any of the other cereal grains [1]. Dunlap et al. work by Chung et al. [61]. This was probably due to a [86,87] reported on corn genotypes with unusual fatty acid smaller variation in the released cultivars used by Ohm compositions (Table 6). They found palmitic acid ranges of and Chung [62]. Samples used by Chung et al. [61] includ-6.3-7.6% and 16.7-18.2% for low and high saturated corn ed some experimental lines. genotypes, respectively. They also reported a range of Bekes et al. [63] investigated 22 hard and 4 soft spring 43.9-46.1% of oleic acids for high oleic acid lines. wheat varieties grown at 3 locations in Canada: varietal Fatty acid composition differs depending on the lipid means ranged from 72 to 134 mg per 10 g (db) flour for extractant (Tables 5 and 6). For example, FL were higher FL, 61-115 mg for NL, 4-11 mg for GL, and 4-9 mg for in both oleic and linoleic acids than the BL of corn and PL (Table 4). There were larger variations in FL contents pearl millet, whereas FL were lower in palmitic acid than for Canadian spring wheats than for U.S. hard winter the BL of millet, oats, and corn. The FA composition of wheats except for GL. Chung [64] showed that U.S. winter NSTL from corn is intermediate to those of FL and BL and spring wheats could not be differentiated by lipid con-based on data complied by Morrison [3]. tents and compositions. Wheat lipid FA compositions for different classes or Unlike the Canadian spring wheats [63], the U.K. soft subclasses are shown in Table 7. The average of 6 HWW winter wheats [65] contained more FL (195-244 mg/10 g, wheats and 14 SWS wheat lipids was lower in palmitic and db) with higher NL content than hard winter wheats stearic acids and higher in linoleic and linolenic acids than (186-210 mg/10 g, db). In general, U.K. hard spring wheats the overall average of 290 wheat lipids. The average FA." In Handbook of Cereal Science and Technology, Revised and Expanded, 435–37. CRC Press, 2000. http://dx.doi.org/10.1201/9781420027228-44.

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Reports on the topic "Tetraploid wheat"

1

Dubcovsky, Jorge, Tzion Fahima, and Ann Blechl. Positional cloning of a gene responsible for high grain protein content in tetraploid wheat. United States Department of Agriculture, September 2003. http://dx.doi.org/10.32747/2003.7695875.bard.

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High Grain Protein Content (GPC) is a desirable trait in breadmaking and pasta wheat varieties because of its positive effects on quality and nutritional value. However, selection for GPC is limited by our poor understanding of the genes involved in the accumulation of protein in the grain. The long-term goal of this project is to provide a better understanding of the genes controlling GPC in wheat. The specific objectives of this project were: a) to develop a high-density genetic map of the GPC gene in tetraploid wheat, b) to construct a T. turgidum Bacterial Artificial Chromosome (BAC) library, c) to construct a physical map of the GPC gene and identify a candidate for the GPC gene. A gene with a large effect on GPC was detected in Triticum turgidum var. dicoccoides and was previously mapped in the short arm of chromosome 6B. To define better the position of the Gpc-B1 locus we developed homozygous recombinant lines with recombination events within the QTL region. Except for the 30-cM region of the QTL these RSLs were isogenic for the rest of the genome minimizing the genetic variability. To minimize the environmental variability the RSLs were characterized using 10 replications in field experiments organized in a Randomized Complete Block Design, which were repeated three times. Using this strategy, we were able to map this QTL as a single Mendelian locus (Gpc-B1) on a 2.6-cM region flanked by RFLP markers Xcdo365 and Xucw67. All three experiments showed that the lines carrying the DIC allele had an average absolute increase in GPC of 14 g/kg. Using the RFLP flanking markers, we established the microcolinearity between a 2.l-cM region including the Gpc-B1 gene in wheat chromosome 6BS and a 350-kb region on rice chromosome 2. Rice genes from this region were used to screen the Triticeae EST collection, and these ESTs were used to saturate the Gpc-B1 region with molecular markers. With these new markers we were able to map the Gpc-B1 locus within a 0.3-cM region flanked by PCR markers Xucw83 and Xucw71. These flanking markers defined a 36-kb colinear region with rice, including one gene that is a potential candidate for the Gpc-B1 gene. To develop a physical map of the Gpc-B1 region in wheat we first constructed a BAC library of tetraploid wheat, from RSL#65 including the high Gpc-B1 allele. We generated half- million clones with an average size of l3l-kb (5.1 X genome equivalents for each of the two genomes). This coverage provides a 99.4% probability of recovering any gene from durum wheat. We used the Gpc-BI flanking markers to screen this BAC library and then completed the physical map by chromosome walking. The physical map included two overlapping BACs covering a region of approximately 250-kb, including two flanking markers and the Gpc-B1 gene. Efforts are underway to sequence these two BACs to determine if additional wheat genes are present in this region. Weare also developing new RSLs to further dissect this region. We developed PCR markers for flanking loci Xucw79andXucw71 to facilitate the introgression of this gene in commercial varieties by marker assisted selection (httQ://maswheat.ucdavis.edu/ orotocols/HGPC/index.hlm). Using these markers we introgressed the Gpc-B1 gene in numerous pasta and common wheat breeding lines.
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2

Dubcovsky, Jorge, Tzion Fahima, Ann Blechl, and Phillip San Miguel. Validation of a candidate gene for increased grain protein content in wheat. United States Department of Agriculture, January 2007. http://dx.doi.org/10.32747/2007.7695857.bard.

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High Grain Protein Content (GPC) of wheat is important for improved nutritional value and industrial quality. However, selection for this trait is limited by our poor understanding of the genes involved in the accumulation of protein in the grain. A gene with a large effect on GPC was detected on the short arm of chromosome 6B in a Triticum turgidum ssp. dicoccoides accession from Israel (DIC, hereafter). During the previous BARD project we constructed a half-million clones Bacterial Artificial Chromosome (BAC) library of tetraploid wheat including the high GPC allele from DIC and mapped the GPC-B1 locus within a 0.3-cM interval. Our long-term goal is to provide a better understanding of the genes controlling grain protein content in wheat. The specific objectives of the current project were to: (1) complete the positional cloning of the GPC-B1 candidate gene; (2) characterize the allelic variation and (3) expression profile of the candidate gene; and (4) validate this gene by using a transgenic RNAi approach to reduce the GPC transcript levels. To achieve these goals we constructed a 245-kb physical map of the GPC-B1 region. Tetraploid and hexaploid wheat lines carrying this 245-kb DIC segment showed delayed senescence and increased GPC and grain micronutrients. The complete sequencing of this region revealed five genes. A high-resolution genetic map, based on approximately 9,000 gametes and new molecular markers enabled us to delimit the GPC-B1 locus to a 7.4-kb region. Complete linkage of the 7.4-kb region with earlier senescence and increase in GPC, Zn, and Fe concentrations in the grain suggested that GPC-B1 is a single gene with multiple pleiotropic effects. The annotation of this 7.4-kb region identified a single gene, encoding a NAC transcription factor, designated as NAM-B1. Allelic variation studies demonstrated that the ancestral wild wheat allele encodes a functional NAC transcription factor whereas modern wheat varieties carry a non-functional NAM-B1 allele. Quantitative PCR showed that transcript levels for the multiple NAMhomologues were low in flag leaves prior to anthesis, after which their levels increased significantly towards grain maturity. Reduction in RNA levels of the multiple NAMhomologues by RNA interference delayed senescence by over three weeks and reduced wheat grain protein, Zn, and Fe content by over 30%. In the transgenic RNAi plants, residual N, Zn and Fe in the dry leaves was significantly higher than in the control plants, confirming a more efficient nutrient remobilization in the presence of higher levels of GPC. The multiple pleiotropic effects of NAM genes suggest a central role for these genes as transcriptional regulators of multiple processes during leaf senescence, including nutrient remobilization to the developing grain. The cloning of GPC-B1 provides a direct link between the regulation of senescence and nutrient remobilization and an entry point to characterize the genes regulating these two processes. This may contribute to their more efficient manipulation in crops and translate into food with enhanced nutritional value. The characterization of the GPC-B1 gene will have a significant impact on wheat production in many regions of the world and will open the door for the identification of additional genes involved in the accumulation of protein in the grain.
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3

Feldman, Moshe, Eitan Millet, Calvin O. Qualset, and Patrick E. McGuire. Mapping and Tagging by DNA Markers of Wild Emmer Alleles that Improve Quantitative Traits in Common Wheat. United States Department of Agriculture, February 2001. http://dx.doi.org/10.32747/2001.7573081.bard.

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The general goal was to identify, map, and tag, with DNA markers, segments of chromosomes of a wild species (wild emmer wheat, the progenitor of cultivated wheat) determining the number, chromosomal locations, interactions, and effects of genes that control quantitative traits when transferred to a cultivated plant (bread wheat). Slight modifications were introduced and not all objectives could be completed within the human and financial resources available, as noted with the specific objectives listed below: 1. To identify the genetic contribution of each of the available wild emmer chromosome-arm substitution lines (CASLs) in the bread wheat cultivar Bethlehem for quantitative traits, including grain yield and its components and grain protein concentration and yield, and the effect of major loci affecting the quality of end-use products. [The quality of end-use products was not analyzed.] 2. To determine the extent and nature of genetic interactions (epistatic effects) between and within homoeologous groups 1 and 7 for the chromosome arms carrying "wild" and "cultivated" alleles as expressed in grain and protein yields and other quantitative traits. [Two experiments were successful, grain protein concentration could not be measured; data are partially analyzed.] 3. To derive recombinant substitution lines (RSLs) for the chromosome arms of homoeologous groups 1 and 7 that were found previously to promote grain and protein yields of cultivated wheat. [The selection of groups 1 and 7 tons based on grain yield in pot experiments. After project began, it was decided also to derive RSLs for the available arms of homoeologous group 4 (4AS and 4BL), based on the apparent importance of chromosome group 4, based on early field trials of the CASLs.] 4. To characterize the RSLs for quantitative traits as in objective 1 and map and tag chromosome segments producing significant effects (quantitative trait loci, QTLs by RFLP markers. [Producing a large population of RSLs for each chromosome arm and mapping them proved more difficult than anticipated, low numbers of RSLs were obtained for two of the chromosome arms.] 5. To construct recombination genetic maps of chromosomes of homoeologous groups 1 and 7 and to compare them to existing maps of wheat and other cereals [Genetic maps are not complete for homoeologous groups 4 and 7.] The rationale for this project is that wild species have characteristics that would be valuable if transferred to a crop plant. We demonstrated the sequence of chromosome manipulations and genetic tests needed to confirm this potential value and enhance transfer. This research has shown that a wild tetraploid species harbors genetic variability for quantitative traits that is interactive and not simply additive when introduced into a common genetic background. Chromosomal segments from several chromosome arms improve yield and protein in wheat but their effect is presumably enhanced when combination of genes from several segments are integrated into a single genotype in order to achieve the benefits of genes from the wild species. The interaction between these genes and those in the recipient species must be accounted for. The results of this study provide a scientific basis for some of the disappointing results that have historically obtained when using wild species as donors for crop improvement and provide a strategy for further successes.
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