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Journal articles on the topic 'Lopingien'

Author: Grafiati

Published: 4 May 2024

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1

Shen, Shu-Zhong, and Yi-Chun Zhang. "Earliest Wuchiapingian (Lopingian, late Permian) brachiopods in southern Hunan, South China: implications for the pre-Lopingian crisis and onset of Lopingian recovery/radiation." Journal of Paleontology 82, no. 5 (September 2008): 924–37. http://dx.doi.org/10.1666/07-118.1.

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The uppermost 5–15 m of the Douling Formation in the southern Hunan area, South China, yields a diverse fauna comprised of ammonoids, bivalves, and brachiopods. The brachiopods reported in this paper consist of 51 species in 34 genera and are dominated by the Lopingian (Late Permian) species associated with a few species persisting from the underlying Maokouan (Late Guadalupian). This fauna is of earliest Wuchiapingian in age as precisely constrained by the associated conodontClarkina postbitteri postbitteriand the Guadalupian-type ammonoid fauna of theRoadoceras-DoulingocerasZone in the brachiopod horizon. The discovery of the Lopingian species-dominated brachiopod fauna in the earliest Wuchiapingian in southern Hunan suggests a much less pronounced effect of the pre-Lopingian crisis (end-Guadalupian mass extinction) than the end-Changhsingian mass extinction in terms of brachiopods, a contemporaneous onset of the Lopingian recovery/radiation during the pre-Lopingian crisis period, and taxonomic selectivity of the pre-Lopingian crisis in terms of different fossil groups. New taxa areEchinauris doulingensisn. sp.,Pararigbyella quadrilobatan. gen. and n. sp. andP. doulingensisn. gen. and n. sp.

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2

Abdala, Fernando, Leandro C. Gaetano, Roger M. H. Smith, and Bruce S. Rubidge. "A new large cynodont from the Late Permian (Lopingian) of the South African Karoo Basin and its phylogenetic significance." Zoological Journal of the Linnean Society 186, no. 4 (May 2, 2019): 983–1005. http://dx.doi.org/10.1093/zoolinnean/zlz004.

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Abstract The Karoo Basin of South Africa has the best global record of Lopingian (Late Permian) non-mammaliaform cynodonts, currently represented by five species. We describe Vetusodon elikhulu gen. et sp. nov., documented by four specimens from the Daptocephalus Assemblage Zone. With a basal skull length of ~18 cm, it is the largest Lopingian cynodont and is also larger than Induan representatives of the group. Vetusodon elikhulu has a cranial morphology that departs notably from that previously documented for Permo-Triassic cynodonts. It features a short and extremely wide snout, resembling that of the contemporaneous therocephalian Moschorhinus, and has large incisors and canines that contrast with the small unicusped postcanines, suggesting a more important role of the anterior dentition for feeding. The dentary is extremely long and robust, with the posterior margin located closer to the craniomandibular joint than in other Lopingian and Induan cynodonts (e.g. Thrinaxodon). The secondary palate morphology of V. elikhulu is unique, being short and incomplete and with the posterior portion of the maxilla partly covering the vomer. A phylogenetic analysis suggests that V. elikhulu is the sister taxon of Eucynodontia and thus the most derived of the Lopingian to Induan cynodonts yet discovered.

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Bond, David P. G., and Paul B. Wignall. "Latitudinal selectivity of foraminifer extinctions during the late Guadalupian crisis." Paleobiology 35, no. 4 (2009): 465–83. http://dx.doi.org/10.1666/0094-8373-35.4.465.

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A global database of middle–upper Permian foraminiferal genera has been compiled from the literature for 75 Guadalupian and 62 Lopingian localities, grouped into 32 and 19 operational geographical units respectively. Cluster analysis reveals that five distinct Guadalupian provinces were reduced to four in the Lopingian, following the disappearance of the Eastern Panthalassa Province. Extinction magnitudes across the Guadalupian/Lopingian (G/L) boundary reveal that, in the remaining provinces, there is a strong regional variation to the losses at low paleolatitudes. The Central and Western Tethys Province experienced a markedly lower extinction magnitude, at both provincial and global levels, than the Eastern and Northern Tethys Province. Panthalassa experienced a high extinction magnitude of endemics, but a global extinction magnitude similar to that recorded in Central and Western Tethys. This regional bias is seen in both the fusulinacean and non-fusulinacean foraminifera, although fusulinaceans suffered much higher magnitudes of extinction. The regional selectivity also persisted during the subsequent Lopingian radiations, with the Central and Western Tethys Province recording the greatest magnitudes. Thus, of 35 new genera recorded globally from the Lopingian, 27 of these are recorded in Central and Western Tethys, compared to five and 12 genera respectively in Panthalassa and in Eastern and Northern Tethys. The Emeishan large igneous province erupted within the Eastern and Northern Tethys Province and may have been a factor in the high extinction–low radiation regime of this region. Regression (and consequent shallow-marine habitat loss) also appears to have been a significant factor. A major, but brief, late Guadalupian regression is best seen in those areas that suffered the greatest extinction losses.

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Xia, W., N. Zhang, Y. Kakuwa, and L. Kakuwa. "Radiolarian and conodont biozonation in the pelagic Guadalupian–Lopingian boundary interval at Dachongling, Guangxi, South China, and mid-upper Permian global correlation." Stratigraphy 2, no. 3 (2005): 217–38. http://dx.doi.org/10.29041/strat.02.3.02.

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High–resolution microbiostratigraphy has identified six conodont zones and five coeval radiolarian zones in the Guadalupian–Lopingian (G-L) transitional interval from a section of middle and upper Permian pelagic cherts at Dachongling, near Qinzhou city in Guangxi, South China. The basal Lopingian GSSP at Penglaitan, South China, is correlated with the base of bed Dch 45-21 at Dachongling on the basis of the first occurrence of the conodont Clarkina postbitteri postbitteri Mei and Wardlaw. The first appearances of the radiolarians Albaillella yamakitai Kuwahara and A. cavitata Kuwahara at the same level indicate that both FADs can be used for identifying theG–L boundary in radiolarian-bearing pelagic cherty facies. The appearance of both radiolarian species in the sections at Sasayama, Gujo-Hachima in Southwest Japan, and in the Quinn River Formation of north-central Nevada, indicates that both boundary-index fossils can be traced worldwide. We therefore suggest that Dachongling secion is a good candidate as an auxiliary reference section for the Guadalupian-Lopingian boundary.

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5

Clapham, Matthew E. "Ecological consequences of the Guadalupian extinction and its role in the brachiopod-mollusk transition." Paleobiology 41, no. 2 (February 24, 2015): 266–79. http://dx.doi.org/10.1017/pab.2014.15.

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AbstractThe Guadalupian (middle Permian) extinction may have triggered substantial ecological restructuring in level-bottom communities, such as turnover in dominant brachiopod genera or a shift from abundant brachiopods to mollusks, despite comparatively minor taxonomic losses. However, ecological changes in relative abundance have been inferred from limited data; as a result, constraints on important shifts like the brachiopod-mollusk transition are imprecise. Here, I reevaluate the magnitude of ecological shifts during the Guadalupian–Lopingian (G-L) interval by supplementing previous census counts of silicified assemblages with counts from non-silicified assemblages and global occurrence data, both sourced from the Paleobiology Database. Brachiopod occurrences are consistent with more pronounced faunal composition changes from the Guadalupian to Lopingian than among stages within those intervals, but only in Iran and South China, and not in Pakistan or a Tethys-wide data set. In Iran and South China, Bray-Curtis dissimilarity values comparing occurrence frequencies between adjacent stages were elevated across the G-L transition, although other intervals exhibited similarly large shifts. However, genus occurrence frequencies were less strongly correlated or were anti-correlated across the G-L transition, suggesting moderate faunal turnover among dominant brachiopod genera. In contrast to previous inferences from silicified faunas, abundances of brachiopods, bivalves, and gastropods remained consistent from the Guadalupian to Lopingian in non-silicified local counts and global occurrences, implying that the brachiopod-mollusk shift did not occur until the end-Permian extinction. Ecological and taxonomic consequences were both minor in level-bottom settings, suggesting that severe environmental perturbations may not be necessary to explain biotic changes during the Guadalupian-Lopingian transition.

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Yuan, Dong-xun, Shu-zhong Shen, and Charles M. Henderson. "Revised Wuchiapingian conodont taxonomy and succession of South China." Journal of Paleontology 91, no. 6 (September 15, 2017): 1199–219. http://dx.doi.org/10.1017/jpa.2017.71.

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AbstractSouth China has become the most important area to establish a global stratigraphic framework of the Wuchiapingian Stage because complete Wuchiapingian sequences include the GSSPs for the base and top of the stage. As the markers of the Wuchiapingian GSSP, conodonts are the most important fossil group to establish the Wuchiapingian biostratigraphic framework. However, few documents have investigated in detail the conodont biostratigraphic succession through the entire Wuchiapingian Stage. Furthermore, the conodont taxonomy of several WuchiapingianClarkinaspecies is still debated. Therefore, we here review all WuchiapingianClarkinaspecies from South China and figure ontogenetic growth series from juvenile to adult individuals for each valid and important species in order to revise both Wuchiapingian conodont taxonomy and the biostratigraphic succession. Based on the Penglaitan, Dukou, and Nanjiang sections, seven conodont zones (Clarkina postbitteri postbitteri,C.dukouensis,C.asymmetrica,C.leveni,C.guangyuanensis,C.transcaucasica, andC.orientalis) are recognized. The WuchiapingianClarkinaspecies lineage is also reviewed to confirm the conodont biostratigraphic framework. The Guadalupian-Lopingian boundary (GLB) interval represents a sequence boundary. The time framework of the pre-Lopingian extinction interval indicates that the beginning of the end-Guadalupian regression is in the upper part of theJinogondolella postserrataZone, and the beginning of the early Lopingian transgression is in the lower part of theClarkina dukouensisZone in South China.

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7

Feng, Yan, Haijun Song, and David P. G. Bond. "Size variations in foraminifers from the early Permian to the Late Triassic: implications for the Guadalupian–Lopingian and the Permian–Triassic mass extinctions." Paleobiology 46, no. 4 (September 30, 2020): 511–32. http://dx.doi.org/10.1017/pab.2020.37.

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AbstractThe final 10 Myr of the Paleozoic saw two of the biggest biological crises in Earth history: the middlePermian extinction (often termed the Guadalupian–Lopingian extinction [GLE]) that was followed 7–8 Myr later by Earth's most catastrophic loss of diversity, the Permian–Triassic mass extinction (PTME). These crises are not only manifest as sharp decreases in biodiversity and—particularly for the PTME—total ecosystem collapse, but they also drove major changes in biological morphological characteristics such as the Lilliput effect. The evolution of test size among different clades of foraminifera during these two extinction events has been less studied. We analyzed a global database of foraminiferal test size (volume) including 20,226 specimens in 464 genera, 98 families, and 9 suborders from 632 publications. Our analyses reveal significant reductions in foraminiferal mean test size across the Guadalupian/Lopingian boundary (GLB) and the Permian/Triassic boundary (PTB), from 8.89 to 7.60 log10 μm3 (lg μm3) and from 7.25 to 5.82 lg μm3, respectively. The decline in test size across the GLB is a function of preferential extinction of genera exhibiting gigantism such as fusulinoidean fusulinids. Other clades show little change in size across the GLB. In contrast, all Lopingian suborders in our analysis (Fusulinina, Lagenina, Miliolina, and Textulariina) experienced a significant decrease in test size across the PTB, mainly due to size-biased extinction and within-lineage change. The PTME was clearly a major catastrophe that affected many groups simultaneously, and the GLE was more selective, perhaps hinting at a subtler, less extreme driver than the later PTME.

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Huttenlocker, Adam K., and Roger M. H. Smith. "New whaitsioids (Therapsida: Therocephalia) from the Teekloof Formation of South Africa and therocephalian diversity during the end-Guadalupian extinction." PeerJ 5 (October 5, 2017): e3868. http://dx.doi.org/10.7717/peerj.3868.

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Two new species of therocephalian therapsids are described from the upper Permian Teekloof Formation of the Karoo Basin, South Africa. They include two specimens of a whaitsiid, Microwhaitsia mendrezi gen. et sp. nov., and a single, small whaitsioid Ophidostoma tatarinovi gen. et sp. nov., which preserves a combination of primitive and apomorphic features. A phylogenetic analysis of 56 therapsid taxa and 136 craniodental and postcranial characters places the new taxa within the monophyletic sister group of baurioids—Whaitsioidea—with Microwhaitsia as a basal whaitsiid and Ophidostoma as an aberrant whaitsioid just outside the hofmeyriid+whaitsiid subclade. The new records support that whaitsioids were diverse during the early-late Permian (Wuchiapingian) and that the dichotomy between whaitsiid-line and baurioid-line eutherocephalians was established early on. The oldest Gondwanan whaitsiid Microwhaitsia and additional records from the lower strata of the Teekloof Formation suggest that whaitsioids had diversified by the early Wuchiapingian and no later than Pristerognathus Assemblage Zone times. Prior extinction estimates based on species counts are reflected in an analysis of origination/extinction rates, which imply increasing faunal turnover from Guadalupian to Lopingian (late Permian) times. The new records support a growing body of evidence that some key Lopingian synapsid clades originated near or prior to the Guadalupian-Lopingian boundary ca. 260–259 million years ago, but only radiated following the end-Guadalupian extinction of dinocephalians and basal therocephalian predators (long-fuse model). Ongoing collecting in older portions of the Teekloof Formation (e.g., Pristerognathus Assemblage Zone) will shed further light on early eutherocephalians during this murky but critical time in their evolutionary diversification.

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9

DAVYDOV, V. I., J. L. CROWLEY, M. D. SCHMITZ, and W. S. SNYDER. "New U–Pb constraints identify the end-Guadalupian and possibly end-Lopingian extinction events conceivably preserved in the passive margin of North America: implication for regional tectonics." Geological Magazine 155, no. 1 (October 25, 2016): 119–31. http://dx.doi.org/10.1017/s0016756816000959.

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AbstractThe discovery and dating of a volcanic ash bed within the upper Phosphoria Formation in SE Idaho, USA, is reported. The ash occurs 11 m below the top of the phosphatic Meade Peak Member and yielded a 206Pb/238U date of 260.57 ± 0.07 / 0.14 / 0.31 Ma, i.e. latest Capitanian, Guadalupian. The stratigraphic position of this ash near the top of the Meade Peak phosphatic Member of Phosphoria Formation indicates plausible completeness of the sedimentation within the Guadalupian–Lopingian and probably at the Permo-Triassic (P-T) transitions. The new radiometric age reveals that the regional biostratigraphy and palaeontology of Phosphoria and Park City formations requires serious reconsideration, particularly in cool water conodonts, bryozoans and brachiopods. The new age proposes that the Guadalupian–Lopingian boundary (GLB) coincides with the Meade Peak – Rex contact and consequently with the end-Guadalupian extinction event. The lack of a major unconformity at the P-T transition suggests that the effects of the Sonoma orogeny were not as extensive as has been assumed.

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Nowak, Hendrik, Elke Schneebeli-Hermann, and Evelyn Kustatscher. "Correlation of Lopingian to Middle Triassic Palynozones." Journal of Earth Science 29, no. 4 (August 2018): 755–77. http://dx.doi.org/10.1007/s12583-018-0790-8.

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11

Shu-Zhong, Shen, G. R. Shi Cao Chang-Qun, Wang Xiang-Dong, and Mei Shi-Long. "Lopingian (Late Permian) stratigraphy, sedimentation and palaeobiogeography in southern Tibet." Newsletters on Stratigraphy 39, no. 2-3 (October 2, 2003): 157–79. http://dx.doi.org/10.1127/nos/39/2003/157.

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12

Vozárová, Anna, Fritz Ebner, Sándor Kovács, Hans-Georg Kräutner, Tibor Szederkenyi, Branislav Krstić, Jasenka Sremac, Dunja Aljinovič, Matevž Novak, and Dragomir Skaberne. "Late Variscan (Carboniferous to Permian) environments in the Circum Pannonian Region." Geologica Carpathica 60, no. 1 (February 1, 2009): 71–104. http://dx.doi.org/10.2478/v10096-009-0002-7.

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Late Variscan (Carboniferous to Permian) environments in the Circum Pannonian RegionThe Pennsylvanian-Cisuralian late-orogenic and post-orogenic paleoenvironments of the Circum Pannonian Region (CPR) include tectono-stratigraphic sequences developed from the Upper Bashkirian-Moscovian marine early molasse stage up to the Guadalupian-Lopingian post-orogenic stage, with gradual connection to the beginning of the Alpine (Neotethyan) sedimentary cycle. Shallow marine siliciclastic or carbonate siliciclastic overstep sequences started in the internal part of the Variscan orogenic belt during the latest Serpukhovian and Bashkirian-Moscovian. They overlapped unconformably the variably metamorphosed Variscan basement, or weakly deformed and metamorphosed foreland and syn-orogenic flysch sediments of Mississippian to Early Pennsylvanian age. The post-Variscan rifting largely affected the Variscan orogenic belt by reactivation of the Variscan lithosphere. The late- to post-orogenic terrestrial sequences started within the internal part of the Variscan orogenic belt during the Middle/Late Pennsylvanian. It continued gradually to terrestrial-shallow water carbonate-siliciclastic sequences in its external part through the Permian. According to the present configuration, the Alpine (Neotethyan) northward shifting transgression started during the Guadalupian/Lopingian in the South and during the Early Triassic in the North.

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MAYS, CHRIS, and STEPHEN MCLOUGHLIN. "END-PERMIAN BURNOUT: THE ROLE OF PERMIAN–TRIASSIC WILDFIRES IN EXTINCTION, CARBON CYCLING, AND ENVIRONMENTAL CHANGE IN EASTERN GONDWANA." PALAIOS 37, no. 6 (June 28, 2022): 292–317. http://dx.doi.org/10.2110/palo.2021.051.

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ABSTRACT Wildfire has been implicated as a potential driver of deforestation and continental biodiversity loss during the end-Permian extinction event (EPE; ∼ 252 Ma). However, it cannot be established whether wildfire activity was anomalous during the EPE without valid pre- and post-EPE baselines. Here, we assess the changes in wildfire activity in the high-latitude lowlands of eastern Gondwana by presenting new long-term, quantitative late Permian (Lopingian) to Early Triassic records of dispersed fossil charcoal and inertinite from sediments of the Sydney Basin, eastern Australia. We also document little-transported fossil charcoal occurrences in middle to late Permian (Guadalupian to Lopingian) permineralized peats of the Lambert Graben, East Antarctica, and Sydney and Bowen basins, eastern Australia, indicating that even vegetation of consistently moist high-latitude settings was prone to regular fire events. Our records show that wildfires were consistently prevalent through the Lopingian, but the EPE demonstrates a clear spike in activity. The relatively low charcoal and inertinite baseline for the Early Triassic is likely due in part to the lower vegetation density, which would have limited fire spread. We review the evidence for middle Permian to Lower Triassic charcoal in the geosphere, and the impacts of wildfires on sedimentation processes and the evolution of landscapes. Moreover, we assess the evidence of continental extinction drivers during the EPE within eastern Australia, and critically evaluate the role of wildfires as a cause and consequence of ecosystem collapse. The initial intensification of the fire regime during the EPE likely played a role in the initial loss of wetland carbon sinks, and contributed to increased greenhouse gas emissions and land and freshwater ecosystem changes. However, we conclude that elevated wildfire frequency was a short-lived phenomenon; recurrent wildfire events were unlikely to be the direct cause of the subsequent long-term absence of peat-forming wetland vegetation, and the associated ‘coal gap' of the Early Triassic.

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Wang, Yu-Jing, Qun Yang, Yen-Nien Cheng, and Jia-Xiang Li. "Lopingian (Upper Permian) radiolarian biostratigraphy of South China." Palaeoworld 15, no. 1 (January 2006): 31–53. http://dx.doi.org/10.1016/j.palwor.2006.03.004.

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15

Zhong, Yu-Ting, Bin He, and Yi-Gang Xu. "Mineralogy and geochemistry of claystones from the Guadalupian–Lopingian boundary at Penglaitan, South China: Insights into the pre-Lopingian geological events." Journal of Asian Earth Sciences 62 (January 2013): 438–62. http://dx.doi.org/10.1016/j.jseaes.2012.10.028.

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Wang, Xiang-Dong, and Xiao-Juan Wang. "Extinction patterns of Late Permian (Lopingian) corals in China." Palaeoworld 16, no. 1-3 (January 2007): 31–38. http://dx.doi.org/10.1016/j.palwor.2007.05.009.

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Shen, Shu-Zhong, Charles M. Henderson, Samuel A. Bowring, Chang-Qun Cao, Yue Wang, Wei Wang, Hua Zhang, Yi-Chun Zhang, and Lin Mu. "High-resolution Lopingian (Late Permian) timescale of South China." Geological Journal 45, no. 2-3 (April 29, 2010): 122–34. http://dx.doi.org/10.1002/gj.1232.

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EDIRISOORIYA, G., H. A. DHARMAGUNAWARDHANE, and STEPHEN MCLOUGHLIN. "The first record of the Permian Glossopteris flora from Sri Lanka: implications for hydrocarbon source rocks in the Mannar Basin." Geological Magazine 155, no. 4 (December 13, 2016): 907–20. http://dx.doi.org/10.1017/s0016756816001114.

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AbstractStrata exposed near Tabbowa Tank, Tabbowa Basin, western Sri Lanka have yielded the first representatives of the distinctive Permian Glossopteris flora from that country. The assemblage includes gymnosperm foliage attributable to Glossopteris raniganjensis, roots referable to Vertebraria australis, seeds assigned to Samaropsis sp., sphenophyte axes (Paracalamites australis) and foliage (Sphenophyllum emarginatum), and fern foliage (Dichotomopteris lindleyi). This small macroflora is interpreted to be of probable Lopingian (late Permian) age based on comparisons with the fossil floras of Peninsula India. Several Glossopteris leaves in the assemblage bear evidence of terrestrial arthropod interactions including hole feeding, margin feeding, possible lamina skeletonization, piercing-and-sucking damage and oviposition scarring. The newly identified onshore Permian strata necessitate re-evaluation of current models explaining the evolution of the adjacent offshore Mannar Basin. Previously considered to have begun subsiding and accumulating sediment during Jurassic time, we propose that the Mannar Basin may have initiated as part of a pan-Gondwanan extensional phase during late Palaeozoic – Triassic time. We interpret the basal, as yet unsampled, seismically reflective strata of this basin to be probable organic-rich continental strata of Lopingian age, equivalent to those recorded in the Tabbowa Basin, and similar to the Permian coal-bearing successions in the rift basins of eastern India and Antarctica. Such continental fossiliferous strata are particularly significant as potential source rocks for recently identified natural gas resources in the Mannar Basin.

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Zhou, Zuren. "Permian basinal ammonoid sequence in Nanpanjiang area of South China—possible overlap between basinal Guadalupian and platform-based Lopingian." Journal of Paleontology 91, S74 (April 2017): 1–95. http://dx.doi.org/10.1017/jpa.2016.128.

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AbstractThe Permian pandemic ammonoids in Nanpanjiang Basin (41 genera, including two new generaGlenisterocerasandFusicrimites, and 56 species, including 21 new species) are systematically described and/or discussed. New species described in this paper areAgathiceras sequaxiliraen. sp.,Akmilleria parahuecoensisn. sp.,Aristoceras liuzhaiensen. sp.,Bamyaniceras nandanensen. sp.,Bamyaniceras yangchangensen. sp.,Bransonoceras longyinensen. sp.,Difuntites furnishin. sp.,Emilites globosusn. sp.,Eoaraxoceras spinosain. sp.,Eumedlicottia kabiensisn. sp.,Fusicrimites nanpanjiangensisn. gen. n. sp.,Glenisteroceras sidazhaiensen. gen. n. sp.,Metaperrinites shaiwaensisn. sp.,Miklukhoceras guizhouensen. sp.,Neocrimites guizhouensisn. sp.,Neopronorites leonovaen. sp.,Popanoceras ziyunensen. sp.,Properrinites gigantusn. sp.,Stacheoceras shaiwaensen. sp.,Svetlanoceras uraloceraformisn. sp., andSynartinskia meyaoensen. sp. A relatively complete Permian basinal ammonoid sequence with six zones has been newly recognized in South China, in ascending order,Properrinites gigantus-Svetlanoceras serpentinum,Svetlanoceras uraloceraformis-Prothalassoceras biforme,Popanoceras kueichowense-Medlicottia orbignyanus,Metaperrinites shaiwaensis-Popanoceras ziyunense,Waagenocerassp.-Propinacoceras beyrichi, andEoaraxoceras spinosai-Difuntites furnishi. The upper three zones are close to being duplicated from the Permian of Las Delicias, Coahuila, Mexico and west Texas, USA; while the lower three zones compare well to those of the Lower Permian in South Urals. TheEoaraxoceras-Difuntitesassemblage, as an index fauna of the upper Capitanian in Coahuila, has been found from the Claystone (3rd) Member of the Shaiwa Formation with the commonly accepted Lopingian stratigraphic age. The updated Permian ammonoid biostratigraphy in South China reveals a possible overlap between the basinal Guadalupian from North America and the platform-based Lopingian from South China.

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Götz, Annette E., P. John Hancox, and Andrew Lloyd. "Permian climate change recorded in palynomorph assemblages of Mozambique (Moatize Basin, eastern Tete Province)." Acta Palaeobotanica 57, no. 1 (June 1, 2017): 3–11. http://dx.doi.org/10.1515/acpa-2017-0001.

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Abstract Knowledge of Late Permian biodiversity patterns, following the end-Guadalupian crisis, is still in its infancy, since most recent studies have focused on the end-Permian biotic crisis. The palynological record of southern Africa, however, reveals major climatic changes during the Late Permian. Here we report new palynological data from eastern Tete Province of Mozambique, documenting the change from cool to warm temperate climates during the Lopingian. This prominent climate signal was also detected recently in other depositional environments elsewhere in southern Africa, and thus enables interregional temporal correlations.

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Shi, Xiao, Jianxin Yu, and Yuewu Sun. "Tyloses in the Lopingian cordaitalean root from Xinjiang, Northwest China." Review of Palaeobotany and Palynology 273 (February 2020): 104134. http://dx.doi.org/10.1016/j.revpalbo.2019.104134.

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22

Shen, S., and G. R. Shi. "Wuchiapingian (early Lopingian, Permian) global brachiopod palaeobiogeography: a quantitative approach." Palaeogeography, Palaeoclimatology, Palaeoecology 162, no. 3-4 (October 2000): 299–318. http://dx.doi.org/10.1016/s0031-0182(00)00133-4.

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23

Shen, Shuzhong, and Neil W. Archbold. "Chonetoidea (Brachiopoda) from the Lopingian (Late Permian) of South China." Alcheringa: An Australasian Journal of Palaeontology 25, no. 3 (January 2001): 327–49. http://dx.doi.org/10.1080/03115510108527806.

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Campi, Monica J., and Guang R. Shi. "New Lopingian (Late Permian) rugosochonetid species from Sichuan, South China." Alcheringa: An Australasian Journal of Palaeontology 29, no. 2 (January 2005): 275–85. http://dx.doi.org/10.1080/03115510508619306.

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GIBSON, MARTHA E. "FIRST REPORT OF FUNGAL PALYNOMORPHS FROM THE ZECHSTEIN GROUP (LOPINGIAN): IMPLICATIONS FOR THE STRATIGRAPHIC COMPLETENESS OF THE EARTH'S PALEOZOIC FUNGAL RECORD." PALAIOS 37, no. 6 (June 28, 2022): 318–29. http://dx.doi.org/10.2110/palo.2021.064.

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ABSTRACT Palynological study of the Permian–Triassic boundary has typically focused on the pollen grain and spore content to reconstruct vegetation, with fungal remains either left unidentified or set aside for future research. Paleozoic fungal microfossil records in particular are lacking. The Zechstein Group (∼ 258–252 Ma; Lopingian) is a remarkable stratigraphic sequence of stacked carbonates and evaporites. High-resolution palynological analysis of new borehole cores through the Zechstein Group of northeast England has revealed its entire sedimentological history and enabled a new reconstruction of vegetation dynamics in central-western Europe preceding the Permian–Triassic boundary. Assemblages composed of conifers, pteridosperms, pteridophytes, sphenopsids, and cycads/ginkgoes were recovered alongside fungal remains throughout the entire sequence. Four fungal morphologies were observed, the most common being smooth-walled spheroidal inclusions of an endobiotic Chytridiomycota or Hypochytridiomycota affinity. Other evidence of fungi includes epiphytic Callimothallus-type fungi (Family Microthyraceae), the dematiaceous Chaetomium-like mold (Family Chaetomiaceae) found associated with soil, cellulose and plant debris, and possible evidence of chytrid-induced pitting on the surface of plant cuticle. This is the first study to highlight the fungal content of Zechstein palynological preparations and while occurrences are rare, they provide new insight into the composition of the Zechstein forest understory, reinforcing the interpretation that the upper Zechstein environment was humid. This work improves our understanding of the taxonomic and functional diversity of fungal taxa associated with evaporite systems during the Lopingian, and highlights the exceptional preservation potential of halite, combating underestimates of fungal richness in the fossil record.

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Xu, Hai-peng, Yi-chun Zhang, Feng Qiao, and Shu-zhong Shen. "A new Changhsingian brachiopod fauna from the Xiala Formation at Tsochen in the central Lhasa Block and its paleogeographical implications." Journal of Paleontology 93, no. 5 (June 14, 2019): 876–98. http://dx.doi.org/10.1017/jpa.2019.28.

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AbstractPermian faunal affinity in the Lhasa Block plays a critical role in reconstructing its paleogeographic evolution. Cisuralian and Guadalupian faunas have been described from the Lhasa Block, but very few Lopingian (late Permian) brachiopods have been reported so far. In this paper, a new diverse brachiopod fauna consisting of 17 species of 17 genera and an unidentifiable Orthotetoidea is described from the uppermost part of the Xiala Formation at the Aduogabu section in the central part of the Lhasa Block. The age of this fauna can be assigned to the Changhsingian (late Lopingian) as indicated by the associated foraminifersColaniella parva(Colani, 1924) andReichelina pulchraMiklukho-Maklay, 1954. Characteristic brachiopods includeSpinomarginifera chengyaoyenensisHuang, 1932,Haydenella wenganensis(Huang, 1932), andAraxathyriscf.dilatatusShen, He, and Zhu, 1992. They also generally suggest a Changhsingian age. Paleobiogeographically, this fauna is uniformly composed of typical Tethyan elements represented bySpinomarginiferaHuang, 1932 andHaydenellaReed, 1944, and some cosmopolitan elements, but no typical cold-water taxa of Gondwanan affinity. This is in contrast to the contemporaneous brachiopod faunas from the Tethys Himalayan region that are characterized by typical cold-water taxa of Gondwanan affinity, e.g.,Costiferina indica(Waagen, 1884),Retimarginifera xizangensisShen et al., 2000,Neospirifer(Quadrospina)tibetensisDing, 1962. Thus, it is strongly indicative that the Lhasa Block had drifted into a relatively warm-water regime during the Changhsingian. An analysis of the paleobiogeographic change of brachiopods in the Lhasa Block throughout the entire Permian further suggests that the Lhasa Block probably had rifted away from the northern peri-Gondwanan margin between the latest Cisuralian and middle Guadalupian, that is, the Neotethys Ocean had opened before middle Guadalupian.

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Gao, Xiangdong, Wuzhong Li, Yiming Yang, Kaigui Yin, and Shihao Zhou. "Multiple Influences of Paleogeography, Sequence Stratigraphy, and Fold Structure on Coalbed Methane Accumulation in the Lopingian (Late Permian) of Junlian Coalfield, SW China." Geofluids 2021 (October 14, 2021): 1–17. http://dx.doi.org/10.1155/2021/3401273.

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Junlian coalfield is one of the main targets for coalbed methane (CBM) exploration and development in the southwest China. Based on field geological survey, core observation, gas content statistics, coal maceral composition, vitrinite reflectance ( R O ), proximate analysis and trace element test, lithological types, lithofacies, sedimentary environment, and structural analysis, this research established the sequence stratigraphy frame, revealed the plane distribution characteristics of sedimentary facies, and defined CBM accumulation mode. The results show that six rock types were identified and further subdivided into twenty lithofacies types. Four types of sedimentary systems such as alluvial plains, delta, lagoon-tidal flat, and carbonate platform were summarized according to their combination characteristics. Additionally, 12-14 fourth-order sequences and three third-order sequences CSI, CSII, and CSIII were divided, and a sequence stratigraphic framework of the Lopingian coal-bearing series was established. Among them, the features of third-order sequence CSIII paleogeography from west to east are alluvial plains, deltas, lagoon-tidal flats, and limited carbonate platforms. Thick coal seams are mainly developed in the sedimentary environment of tidal flats, delta plains, and floodplains behind banks. Closely related to coal seam thickness, gas contents of Lopingian coal seams are generally higher than 8 m3/t, except the low level in northwest and partial denudation areas. CBM accumulation is significantly controlled by the fold structure, and the hydraulic plugging effect makes the syncline core favorable for CBM accumulation. Furthermore, favorable geostress conditions enable the secondary anticline to become a favorable area for CBM accumulation when the sealing conditions are better. This research will provide a theoretical guide for the exploration and development of CBM in the study area.

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Yang, Ji-Yuan, Hai-Bo Wei, Xu-Dong Gou, Shi-Ling Yang, and Zhuo Feng. "Leaf anatomy of Ningxiaites specialis from the Lopingian of Northwest China." Review of Palaeobotany and Palynology 300 (May 2022): 104632. http://dx.doi.org/10.1016/j.revpalbo.2022.104632.

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Ramsdale, Rick. "A possible serpulid tube worm of the genus Filograna from the upper Permian Cadeby Formation of South Yorkshire, UK." Proceedings of the Yorkshire Geological Society 63, no. 3 (January 12, 2021): pygs2020–016. http://dx.doi.org/10.1144/pygs2020-016.

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Simple, tubular clusters from a single boundstone hand specimen, collected from loose material in the Permian (Lopingian) Cadeby Formation at Hazel Lane Quarry, Doncaster Metropolitan Borough, are tentatively identified as the serpulid worm, Filograna sp. A. The tubes are calcareous, found in vertical clusters, with some short sections growing horizontally, and with external diameters between 0.6 and 1 mm. Recent work on the Permian Tethyan deposits of Italy have identified occurrences of the genus Filograna from before the Permian–Triassic extinction, but the importance of this example is its provenance in the Zechstein English Shelf biota, where reported occurrences of Serpulidae have been rare.

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30

Maldonado, Amy L., and Paula J. Noble. "Radiolarians from the upper Guadalupian (Middle Permian) Reef Trail Member of the Bell Canyon Formation, West Texas and their biostratigraphic implications." Micropaleontology 56, no. 1-2 (2010): 69–115. http://dx.doi.org/10.47894/mpal.56.1.03.

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Moderately well-preserved radiolarians are described from two measured sections of the Reef Trail Member of the Bell Canyon Formation (Middle Permian), exposed in the Patterson Hills, Guadalupe Mountains National Park, West Texas. The Reef Trail Member is the youngest member of a series of basinal marine carbonates deposited in the Delaware basin prior to the deposition of the Castile Formation evaporites, and represents the uppermost Guadalupian (upper Capitanian). The Guadalupian-Lopingian boundary is presumed to occur at or just above the top of the Reef Trail-Castile contact because of the presence of C. postbitteri hongshuiensis ~ 2m below the top of the Reef Trail Member in sections in the Patterson Hills. A total of 30 genera containing 51 species were recovered from the Reef Trail Member, belonging to the orders Albaillellaria, Latentifistularia, Spumellaria, and Entactinaria. The following new taxa are described: Raphidociclicus scutum, Camptoalatus volaticus, Pseudoalbaillella delawarensis, Astroentactinia porosa, Stigmosphaerostylus favusa, Polyedroentactinia quadrata, Praedeflandrella firmata, P. prolata, Copicyntra spinosa, Copicyntra irregulata, and Paracopicyntra puncta, and Copicyntroides nazarovi. The following taxa are emended: Raciditor scalae (Caridroit and De Wever 1986), and Klaengspongus Sashida 2000b. Near the top of the Reef Trail Member, there are marked fluctuations in the relative abundances of albaillellarian and latentifistularian species. Fluctuations are possibly controlled by paleoenvironmental changes associated with the onset of the end-Guadalupian extinction event. Biostratigraphically, the fauna most closely correlates with the widely recognized F. bipartitus – F. charveti Zone sensu Caridroit, which we now consider to be uppermost Guadalupian, not Lopingian, and to the Guadalupian F. charveti Zone of Sun and Xia 2006 from oceanic facies in China. Both older and younger biostratigraphic markers also occur in the Reef Trail fauna, including Pseudoalbaillella longtanensis and A. yamakitai, requiring a reevaluation of the reliability of these albaillellarian taxa in biostratigraphy.

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AREFIFARD, SAKINEH. "Guadalupian cool versus warm water deposits in central Iran: a record of the Capitanian Kamura event." Geological Magazine 156, no. 3 (October 26, 2017): 430–46. http://dx.doi.org/10.1017/s0016756817000863.

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AbstractAn integration of geochemical and grain association studies were carried out on Middle Permian deposits in central Iran where both cool and warm water carbonates are found. The recrystallization of most bioclasts, lime-mud matrix and ooids along with high Sr contents suggests a probable original aragonite mineralogy for carbonates of the Middle Permian Jamal Formation at the Shotori section. Low bulk carbonate δ18O values imply pervasive diagenetic alteration in this section. Conversely, Middle Permian deposits at the correlative Bagh-e Vang section have a probable calcite precursor supported by low Sr contents and no evidence of recrystallization. This mineralogical variation in these coeval carbonates is considered to be due to the change in depth and temperature of the depositional palaeoenvironment. δ13C values started to rise over 2 ‰ PDB and reached a maximum of 4.3 ‰ PDB at the Wordian–Capitanian boundary at the Bagh-e Vang section. This δ13C rise is attributed to high primary productivity as previously reported in the Capitanian Abadeh Formation in central Iran. The positive δ13C excursion in these sections is correlated with the Capitanian ‘Kamura event’ identified from the mid-Panthalassian sections in Japan. No noticeable positive excursion occurs in the δ13C plot at the Shotori section making the interpretation of palaeo-productivity difficult. It is suggested that an active oceanic upwelling was the probable driver of the Middle Permian oceanic productivity in central Iran. Remarkable negative δ13C excursions around 3.7 and 4.2 ‰ PDB in Capitanian carbonates close to the Guadalupian–Lopingian boundary at the Bagh-e Vang and Abadeh sections, respectively are recorded, which are a proxy for low palaeo-productivity and a transition from a cool to warm climate, consistent with an early Lopingian sea level rise.

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Shen, Shu-Zhong, and G. R. Shi. "Latest Guadalupian brachiopods from the Guadalupian/Lopingian boundary GSSP section at Penglaitan in Laibin, Guangxi, South China and implications for the timing of the pre-Lopingian crisis." Palaeoworld 18, no. 2-3 (September 2009): 152–61. http://dx.doi.org/10.1016/j.palwor.2009.04.010.

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33

Vachard, Daniel, Jérémie Gaillot, Denis Vaslet, and Yves-Michel Le Nindre. "Foraminifers and algae from the Khuff Formation (late Middle Permian-Early Triassic) of central Saudi Arabia." GeoArabia 10, no. 4 (October 1, 2005): 137–86. http://dx.doi.org/10.2113/geoarabia1004137.

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ABSTRACT Algae and smaller foraminifers of the eponymous Khuff Formation (Saudi Arabia) principally comprise Permocalculus, biseriamminids, hemigordiids and lagenids. Due to the end-Capitanian crisis (Late/Middle Permian boundary) and the regional palaeoecology, fusulinids are rare and only represented by Nankinella sp. and Eostaffella? sp. Palaeofusulinids are completely lacking. New age data shows that these foraminifers correspond to the complete Lopingian (Late Permian) as indicated by several species of Paradagmarita. The position of the Triassic/Permian Boundary is approximately characterised, but requires more accurate studies. Forty-three taxa were identified, mostly in open nomenclature. One new species is described: Glomospirella? linae n. sp. The foraminiferal assemblage is correlated with several associations in Iran, Turkey, Transcaucasia and south China.

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Phillips, L. J., J. S. Esterle, and S. A. Edwards. "Review of Lopingian (upper Permian) stratigraphy of the Galilee Basin, Queensland, Australia." Australian Journal of Earth Sciences 64, no. 3 (March 7, 2017): 283–300. http://dx.doi.org/10.1080/08120099.2017.1290684.

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35

Dong, Yixin, Shenglin Xu, Long Wen, Hongde Chen, Siyi Fu, Yijiang Zhong, Jiuyuan Wang, Peng Zhu, and Ying Cui. "Tectonic control of Guadalupian-Lopingian cherts in northwestern Sichuan Basin, South China." Palaeogeography, Palaeoclimatology, Palaeoecology 557 (November 2020): 109915. http://dx.doi.org/10.1016/j.palaeo.2020.109915.

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36

Wang, Wei, Changqun Cao, and Yue Wang. "The carbon isotope excursion on GSSP candidate section of Lopingian–Guadalupian boundary." Earth and Planetary Science Letters 220, no. 1-2 (March 2004): 57–67. http://dx.doi.org/10.1016/s0012-821x(04)00033-0.

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37

Shi, G. R., J. B. Waterhouse, and S. McLoughlin. "The Lopingian of Australasia: a review of biostratigraphy, correlations, palaeogeography and palaeobiogeography." Geological Journal 45, no. 2-3 (March 22, 2010): 230–63. http://dx.doi.org/10.1002/gj.1213.

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Waterhouse, J. B. "Lopingian (Late Permian) stratigraphy of the Salt Range, Pakistan and Himalayan region." Geological Journal 45, no. 2-3 (April 29, 2010): 264–84. http://dx.doi.org/10.1002/gj.1235.

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39

SHEN, SHU-ZHONG, and MATTHEW E. CLAPHAM. "WUCHIAPINGIAN (LOPINGIAN, LATE PERMIAN) BRACHIOPODS FROM THE EPISKOPI FORMATION OF HYDRA ISLAND, GREECE." Palaeontology 52, no. 4 (July 2009): 713–43. http://dx.doi.org/10.1111/j.1475-4983.2009.00879.x.

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40

Ye, FaCheng, XinChun Liu, Wei Wang, XiaoZheng Chen, Jing Liu, ShuZhong Shen, WenQian Wang, et al. "Biostratigraphy constraining strontium isotopic stratigraphy and its application on the Lopingian (Late Permian)." Science China Earth Sciences 58, no. 11 (July 2, 2015): 1951–59. http://dx.doi.org/10.1007/s11430-015-5134-2.

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41

Shen, Shu-Zhong, Charles M. Henderson, and Ian D. Somerville. "Preface: Lopingian (Late Permian) stratigraphy of the world, major events and environmental change." Geological Journal 45, no. 2-3 (April 5, 2010): 119–21. http://dx.doi.org/10.1002/gj.1239.

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42

Pritchard, Adam C., Hans-Dieter Sues, Diane Scott, and Robert R. Reisz. "Osteology, relationships and functional morphology of Weigeltisaurus jaekeli (Diapsida, Weigeltisauridae) based on a complete skeleton from the Upper Permian Kupferschiefer of Germany." PeerJ 9 (May 20, 2021): e11413. http://dx.doi.org/10.7717/peerj.11413.

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Background Weigeltisauridae is a clade of small-bodied diapsids characterized by a horned cranial frill, slender trunk and limbs, and a patagium supported by elongated bony rods. Partial skeletons and fragments are definitively known only from upper Permian (Lopingian) rocks in England, Germany, Madagascar and Russia. Despite these discoveries, there have been few detailed descriptions of weigeltisaurid skeletons, and the homologies of many skeletal elements—especially the rods supporting the patagium—remain the subject of controversy. Materials & Methods Here, we provide a detailed description of a nearly complete skeleton of Weigeltisaurus jaekeli from the upper Permian (Lopingian: Wuchiapingian) Kupferschiefer of Lower Saxony, Germany. Briefly addressed by past authors, the skeleton preserves a nearly complete skull, postcranial axial skeleton, appendicular skeleton, and patagial supports. Through comparisons with extant and fossil diapsids, we examine the hypotheses for the homologies of the patagial rods. To examine the phylogenetic position of Weigeltisauridae and characterize the morphology of the clade, we integrate the material and other weigeltisaurids into a parsimony-based phylogenetic analysis focused on Permo-Triassic non-saurian Diapsida and early Sauria (61 taxa, 339 characters). Results We recognize a number of intriguing anatomical features in the weigeltisaurid skeleton described here, including hollow horns on the post-temporal arch, lanceolate teeth in the posterior portion of the maxilla, the absence of a bony arch connecting the postorbital and squamosal bones, elongate and slender phalanges that resemble those of extant arboreal squamates, and patagial rods that are positioned superficial to the lateral one third of the gastral basket. Our phylogenetic study recovers a monophyletic Weigeltisauridae including Coelurosauravus elivensis, Weigeltisaurus jaekeli, and Rautiania spp. The clade is recovered as the sister taxon to Drepanosauromorpha outside of Sauria (=Lepidosauria + Archosauria). Conclusions Our anatomical observations and phylogenetic analysis show variety of plesiomorphic diapsid characters and apomorphies of Weigeltisauridae in the specimen described here. We corroborate the hypothesis that the patagial ossifications are dermal bones unrelated to the axial skeleton. The gliding apparatus of weigeltisaurids was constructed from dermal elements unknown in other known gliding diapsids. SMNK-PAL 2882 and other weigeltisaurid specimens highlight the high morphological disparity of Paleozoic diapsids already prior to their radiation in the early Mesozoic.

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43

Zou, Jianhua, Feng Han, Tian Li, Heming Tian, and Yingjiao Li. "Mineralogical and Geochemical Compositions of the Lopingian Coals in the Zhongliangshan Coalfield, Southwestern China." Minerals 8, no. 3 (March 6, 2018): 104. http://dx.doi.org/10.3390/min8030104.

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44

Wang, Y., and K. Ueno. "A NEW FUSULINOIDEAN GENUS DILATOFUSULINA FROM THE LOPINGIAN (UPPER PERMIAN) OF SOUTHERN TIBET, CHINA." Journal of Foraminiferal Research 39, no. 1 (January 1, 2009): 56–65. http://dx.doi.org/10.2113/gsjfr.39.1.56.

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45

Rasnitsyn, A. P., and D. E. van Dijk. "The First GondwananEpimastaxfrom the Lopingian of KwaZulu-Natal, South Africa (Insecta: Palaeomanteida = Miomoptera: Permosialidae)." African Invertebrates 52, no. 1 (June 2011): 207–9. http://dx.doi.org/10.5733/afin.052.0110.

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46

Zhang, Maochao, Zheng Gong, Yu Zhou, Yanmei Liu, Jun Li, and Chengmin Huang. "Cooler Equatorial Climate in the Late Lopingian Estimated from Paleosols Developed on Emeishan Basalts." Journal of Geology 130, no. 1 (January 1, 2022): 23–44. http://dx.doi.org/10.1086/718351.

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47

Zhou, Yu, Yun Guo, Josef Pšenička, Jiří Bek, Shi-Ling Yang, and Zhuo Feng. "A new marattialean fern, Pectinangium xuanweiense sp. nov., from the Lopingian of Southwest China." Review of Palaeobotany and Palynology 295 (December 2021): 104500. http://dx.doi.org/10.1016/j.revpalbo.2021.104500.

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48

Phillips, L. J., S. A. Edwards, V. Bianchi, and J. S. Esterle. "Paleo-environmental reconstruction of Lopingian (upper Permian) sediments in the Galilee Basin, Queensland, Australia." Australian Journal of Earth Sciences 64, no. 5 (June 25, 2017): 587–609. http://dx.doi.org/10.1080/08120099.2017.1338618.

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49

Wang, Shi-Jun, Xiao-Yuan He, and Long-Yi Shao. "Cycad Wood from the Lopingian (Late Permian) of Southern China:Shuichengoxylon tianiigen. et sp. nov." International Journal of Plant Sciences 172, no. 5 (June 2011): 725–34. http://dx.doi.org/10.1086/659454.

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50

Kossovaya, Olga L., and Dieter Weyer. "Lopingian corals from the Omolon Massif (Eastern Siberia), the northernmost Permian boreal Rugosa community." Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 287, no. 2 (February 1, 2018): 167–94. http://dx.doi.org/10.1127/njgpa/2018/0711.

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