Dissertations / Theses on the topic 'Ecological disturbance'

Thesis (Ph. D.)--University of Florida, 1999.
Title from first page of PDF file. Document formatted into pages; contains ix, 261 p.; also includes graphics. Vita. Includes bibliographical references (p. 248-260).

Thesis (Master of Community Planning)--University of Cincinnati, 2009.
Advisor: Menelaos Triantafillou. Title from electronic thesis title page (viewed Aug. 18, 2009). Includes abstract. Keywords: Ecological Restoration; Urban Planning; Integration; Landscape Ecology. Includes bibliographical references.

Interactions among site conditions, disturbance events, and climate determine the patterns of forest species recruitment and mortality across landscapes. Forests of the American Southwest have undergone significant changes over a century of altered disturbance regimes, human land uses, and changing environmental conditions. This study reconstructs the interactions between fire, spruce beetle outbreaks, climate, and anthropogenic factors and their influence on the species composition, spatial extent, and structure of four upper elevation forest types. We found that fire-climate associations changed following fire exclusion and recent high-severity fires occurred during less severe conditions than in several larger, lower severity fires in the historical record. Contemporary fires are burning with higher severity than similarly-sized historical fires, suggesting a shift toward higher-severity fire as a result of changes to forest structure and fuels over much of the upper elevation forest. In high elevation forests, the area occupied by Engelmann spruce and corkbark fir doubled in size over the four decades following fire exclusion. The increase in spruce beetle outbreak size and severity in the 20th century appears to be linked to significant expansion of host extent, accelerated growth of spruce in mixed-conifer forest, and incidence of anomalously warm summer temperatures followed by up to a decade of low precipitation. Trends toward warming, drying conditions are expected to increase the risk of future high-severity outbreaks, especially in locations of recent spruce population expansion. Forest conversion from disturbance-adapted to competition-adapted species following fire exclusion was a function of site productivity. Species assemblages in the lowest and highest productivity sites were the most stable over the century following fire exclusion. Frequent low severity fires maintained the stocking of forests in moderate productivity sites below their biological potential, conferring a degree of resistance to drought, insect outbreaks, and high-severity fire prior to fire exclusion. Current forests located on moderate productivity sites are now the most vulnerable to drought and future disturbance. Aggressive action to restore historical species composition, stocking and fire component of these forests may return resilience to this system in the face of projected changes to fire and climate dynamics.

This thesis set out to investigate the influence of disturbance on the succession of the Krakatau islands (Rakata, Sertung, Panjang). The hierarchical model of succession by S. Pickett and colleagues (1987) was adopted as a research framework, and provided the basis for an alternative model of succession on Krakatau that focuses on processes rather than successional pathways. Investigations were conducted on (i) the meso-scale, and (ii) the patch-scale, (i) quantified the recent disturbance regime, and inter- and intra-island differences in diversity, (ii) compared sapling performance (growth, mortality and recruitment), and species compositional patterning in space and time for saplings and the seed bank with respect to island, gap size and severity of disturbance. Multivariate techniques were used, and amongst other attempts at characterising the light environment, hemispherical photography was employed. For the first time the effect of a continuous period of volcanic activity (1992-1995) of Anak Krakatau could be directly quantified and compared between Panjang and Sertung (ash-affected) and Rakata (receiving no ash). Increased rates of gap formation in the volcanically active period in comparison to the previous decade were found for all islands. This supports the disturbance-driven model of Whittaker and colleagues. However, an extension is required, because, contrary to expectation, Rakata also experienced more disturbance. This increase is argued to be a result of more severe weather conditions, and an increased number of earth tremors, during times of volcanic activity. The disturbance factors of extreme climatic events (e.g. ENSO events) and human impact are also proposed for inclusion in the alternative model. Drought associated with the 1994 El Niño is of relevance to short-term and potential long-term impact on regeneration dynamics and succession. Attention was drawn to the local human influence of pumice mining on the coastal forests. Supporting previous findings on the plot- and whole island scales, data from species presence/absence transects established that species richness and beta-diversity on the ash- affected islands was also lower on the meso-scale. Panjang's canopy composition is less uniform, and locally more species-rich than Sertung's. More evidence of the suggested decline of the mono-dominant species Neonauclea calycina and Timonius compressicaulis was gathered. The third dominant, Dysoxylum gaudichaudianum, is expanding in the lowlands of all islands. This is aided by its ability to regenerate in moderate shade, to grow rapidly in gap environments, and its tolerance of ash-fall, drought and herbivory. However, on Rakata, it is not expected to become generally mono-dominant because a considerable number of other potential canopy species are present. Sapling performance and species composition and its changes were in general strongly affected by ash-fall and drought. These factors tended to override effects of gap size and severity of disturbance. Advance regeneration, and the composition of the local forest type were identified as important factors influencing the composition of the early stages of gap-fill. The local forest type also seemed to contribute most to seed bank composition. As rarer species tended to have clumped distributions, and 'safe sites' for regeneration seemed not to be limiting, dispersal constraints were argued to be the most likely factors slowing diversification, unless further severe volcanic disturbance leads to successional set-back. The latter also strongly limits the predictability of succession on Krakatau.

Thesis (Ph. D.)--University of Missouri-Columbia, 2005.
The entire dissertation/thesis text is included in the research.pdf file; the official abstract appears in the short.pdf file (which also appears in the research.pdf); a non-technical general description, or public abstract, appears in the public.pdf file. Title from title screen of research.pdf file viewed on (November 14, 2006) Vita. Includes bibliographical references.

The Arabidopsis HapMap project, and follow-on work carried out by the Bergelson and Nordborg groups, established in broad outline the demographic history and population structure of wild Arabidopsis thaliana. Genome‐wide association studies are likewise making considerable advances in identifying genes associated with ecologically significant traits, and thus in identifying candidate genes likely to be under the action of natural selection. The aim of this project has been to further expand and combine these lines of investigation, by using genomic data to test ecological hypotheses and to grant more complete insight into the rangeof selection pressures acting upon wildpopulations. A method to measure and elucidate the genetic similarity of genomic regions between sampled accessions was therefore developed to facilitate this. 250K SNP data from RegMap accessions was then examined for evidence of patterns of migration and gene flow across Europe. Those observations formed the basis of a simple model of the history of the UK population relative to that of Europe. Comparisons of observed genotypes against expectations derived from the modelallowed the identification of genomic regions under the influence of selection. Loci corresponding to signatures of selection indicated positive selection acting upon phenotypes of disease resistance, flowering time, and seed size.

Thesis (PhD)--Stellenbosch University, 2014.
ENGLISH ABSTRACT: Globally, habitat transformation causes biodiversity loss, with the transformed matrix often affecting the disturbance regime in remnant natural patches. In South Africa, significant parts of the Indian Ocean coastal belt and grassland biomes have been transformed into commercial forestry plantations of alien trees, which are detrimental to local biodiversity. Consequently, large scale ecological networks (ENs) of remnant natural vegetation, maintained areas (e.g. firebreaks) and special landscape features (e.g. rocky outcrops and wetlands) have been implemented among forestry compartments to offset the negative effect of this land use on biodiversity. Different grassland areas, which constitute a major portion of ENs, were managed in different ways, as governed by their primary purpose (e.g. fire protection or conservation). The overall aim of this study was to determine how grassland floral and grasshopper herbivore communities responded to different disturbances (mowing, burning and grazing), and how we can adjust management of the major disturbances to effectively conserve these major components of biodiversity in ENs. Sampling was carried out in the commercial forestry ENs in the lower-elevation Zululand area and adjacent reserve area iSimangaliso Wetland Park, as well as in the forestry ENs in the higher-lying Midlands and adjacent iMpendle Nature Reserve. Both the reserves or protected areas (PAs) acted as reference sites, while other sites were chosen to represent the predominant disturbances in ENs at each locality: mowing, annual vs. longer-rotation burning, time since last fire, and domestic cattle grazing. In the Zululand subtropical grassland (chapter 2), I explored the effect of frequent mowing on firebreaks, and the effect of patch size and isolation on plant communities in non-firebreak natural areas of the EN. Frequent mowing resulted in plant species loss and a shift in species composition of firebreaks. Furthermore, small, isolated patches in the EN far away from the PA border had lower plant species richness and greater species turnover than wide, interconnected corridors near the PA border, which, in turn, was similar to reference sites in the PA. As plant species were lost from frequently-mown firebreaks and small, isolated patches in the EN, I recommend that this management practice should be confined to demarcated areas (e.g. forestry compartment edges and firebreaks) and that creation of wide, interconnected corridors should be prioritized when designing ENs. In higher elevation Afromontane grassland (chapter 3), I investigated the effect of annual burning on plant communities in firebreaks by comparing them to less frequently burned grassland in the EN and PA, respectively. Grazing by domestic cattle was taken as an embedded factor for firebreak and less frequently burned sites in the EN. There were three firebreak types: annually-burned with heavy cattle grazing (plantation firebreaks), annually-burned with light cattle grazing (peripheral firebreaks), and annual burning without cattle grazing (PA firebreaks). Burned reference grassland in the EN and PA hosted plant communities that were similar in species richness, composition and turnover. This was also the case for lightly-grazed peripheral EN firebreaks and PA firebreaks. However, species composition and turnover of plantation EN firebreaks with heavy cattle grazing differed from that in the other two firebreak types. Although not significant (P<0.1), plantation EN firebreaks had less plant species than reference burned grassland in the EN, and all firebreak types had less plant species, lower species turnover and different species composition when compared to reference burned grassland in the PA. Annual burning of firebreaks, with and without cattle grazing, caused a significant shift in plant species composition and a reduction in plant species turnover. When annual burning was combined with heavy cattle grazing, plant species were lost, as was the case in plantation EN firebreaks. Therefore, I recommend that this management practice should be confined to firebreaks, and that cattle access to firebreaks should be strictly controlled. In Chapter 4, I considered the effect of cattle grazing (presence vs. absence, as well as intensity) on Afromontane grassland against the natural backdrop of variation caused by time since last fire in grassland with longer fire-return intervals (excluding all firebreaks). Lowest plant species richness and turnover occurred in unburned (i.e. burned >12 months prior to sampling), ungrazed grassland in the PA. Burning and grazing both caused a change in plant species composition that went hand in hand with an increase in plant species richness and turnover. However, burning (burned vs. unburned) only affected plant communities in ungrazed grassland in the PA. Similarly, the presence of large mammalian grazers (EN vs. PA) only affected plant communities in unburned grassland. Unburned plant communities grazed by domestic cattle in the EN were similar to those in the PA grazed by indigenous black wildebeest, indicating that cattle grazing simulates, at least to some degree, the effect of indigenous ungulate grazing. Nevertheless, heavily-grazed grassland had less plant species than moderately-grazed grassland in the EN. I recommend that burning and grazing should continue in grassland ENs, as these natural disturbances are necessary to maintain diverse and dynamic ecosystems. Nevertheless, managers should instigate cattle grazing with caution, as high intensity grazing can be detrimental to conservation efforts. In Chapter 5, I examined the effect of annual burning, cattle grazing (presence vs. absence) and time since last fire on grasshopper assemblages in Afromontane grassland. In general, grasshoppers benefitted from disturbance, and were remarkably resilient to different disturbance regimes. Grasshopper species richness and their abundance were both greatest in annually-burned firebreaks with light cattle grazing, and lowest in moribund grassland in the PA which had not been burned for several years. Yet, time since last fire only affected grasshopper communities in the absence of large grazers (in the PA). None of the individual disturbances had an effect on the grasshopper assemblage. Rather, these insects responded to the combined effect of annual burning with cattle grazing. Sites were similar in grasshopper species richness, composition and abundance whenever either annual burning or cattle were absent, which suggests that these two disturbances drive changes in the grasshopper assemblage in these grasslands. Although grasshoppers benefited from annual burning with light cattle grazing, I would not recommend this disturbance regime outside firebreaks. Rather, management of other grassland areas in the EN should adapt longer fire-return intervals with a rotational cattle grazing system, so that undisturbed habitat is provided for other sensitive taxa. In conclusion, grassland plants and grasshoppers benefited from some form of disturbance, but were lost from small, isolated patches in the EN, as well as from areas with high disturbance frequency and intensity. Simulation of natural disturbances (moderate levels of fire and grazing) in wide, interconnected corridors is necessary for maintaining diverse and dynamic grassland ecosystem in ENs among commercial forestry plantations.
AFRIKAANSE OPSOMMING: Omskepping van natuurlike habitat na lande of plantasies veroorsaak biodiversiteitsverlies wêreldwyd. Boonop het sulke veranderinge dikwels 'n effek op die versteurings binne-in oorblywende kolle natuurlike plantegroei wat verreikende gevolge kan hê. Groot gedeeltes van die Suid-Afrikaanse grasveldbioom is omskep in bosbou plantasies wat bestaan uit uitheemse bome wat 'n baie groot nadelige effek op plaaslike biodiversiteit het. Daarom is grootskaalse ekologiese netwerke (EN’e), wat bestaan uit oorblywende kolle natuurlike plantegroei, brandbane en spesiale habitattipes in die landskap (bv. rotsriwwe en vleilande), tussen bosbouplantasies geïmplimenteer met die doel om die negatiewe effek van plantasies op plaaslike biodiversiteit te verlig. Bestuur van grasvelde, wat 'n groot gedeel van EN’e uitmaak, wissel dikwels en hang af van hulle primêre doel (bv. beskerming van plantasies teen wegholveldbrande of natuurbewaring). Die doel van hierdie projek was om vas te stel hoe plant- en springkaangemeenskappe in grasvelde reageer op verskillende versteurings (grassny, brand en beweiding), en die optimale bestuur van die versteurings om die biodiversiteit in grasvelde beter te bewaar. Steekproewe is geneem in EN’e tussen bosbouplantasies in die laagliggende Zululand en langsliggende wêrelderfenisgebied, iSimangaliso Wetland Park, asook in die hoërliggende Midlands en langsliggende iMpendle Natuurreservaat (NR). NR’e het as verwysing gedien waarteen die effek van grassny, frekwensie van brande, tydsverloop vanaf die laaste brand, en beweiding deur beeste, wat tipiese versteuringe in EN’e is, gemeet is. In hoofstuk 2 het ek vasgestel wat die effek van grassnyfrekwensie op plantgemeenskappe in brandbane is, en hoe plantgemeenskappe in subtropiese grasveld in die res van die EN reageer op die grootte en strukturele isolasie van oorblywende kolle natuurlike plantegroei. 'n Hoë grassnyfrekwensie het 'n verandering in die spesiesamestelling van plantgemeenskappe in brandbane veroorsaak wat gepaard gegaan het met spesiesverlies. Terselfdertyd was daar minder plant spesies in klein, geïsoleerde kolle natuurlike plantegroei as wat daar in wyer, aaneenskakelende gange nader aan die natuurreservaatgrens was. Laasgenoemde het plantgemeenskappe bevat wat baie soortgelyk aan die in die natuurreservaat was. Daarom stel ek voor dat die skep van wye, aaneengeskakelde natuurlike habitat prioriteit moet geniet wanneer nuwe EN’e ontwerp word, en dat gras slegs gereeld gesny moet word in spesifieke, afgebakende areas (bv. brandbane). Die rede hiervoor is dat hierdie bestuurspraktyk nie bevorderlik was vir die bewaring van plantdiversiteit in EN’e nie. In hoofstuk 3 het ek gekyk hoe die plantgemeenskappe in brandbane daarop reageer om elke jaar gebrand te word deur hulle te vergelyk met Afrikaberg grasveld in die EN en NR wat minder gereeld gebrand word. Beweiding deur beeste is gesien as 'n integrale deel van die EN. Ek het onderskei tussen plantasiebrandbane met swaar beweiding, randbrandbane met ligte beweiding en brandbane in die NR sonder beweiding. Die plantspesiesamestelling van brandbane, met ligte of geen beweiding nie, het verskil van grasvelde wat minder gereeld gebrand word. Tog is die hoeveelheid plantspesies nie geraak nie. Alhoewel die plantgemeenskappe in ligbeweide brandbane soos die in onbeweide brandbane in die NR was, het die plantspesiesamestelling van beide verskille getoon wanneer hulle vergelyk is met plantasiebrandbane wat swaarder deur beeste bewei is. Plantspesierykheid in plantasiebrandbane was boonop heelwat laer as wat in NR grasvelde gevind is, en daar was heelwat meer kaal grond in plantasiebrandbane as in enige van die ander areas. Oor die algemeen het plantspesiesrykheid van brandbane nie daaronder gely om elke jaar gebrand te word nie, maar kwesbare plantgemeenskappe in brandbane het wel daaronder gely om swaar bewei te word. Daarom stel ek voor dat jaarlikse brande tot brandbane beperk word en dat beeste se toegang tot brandbane streng beheer word. In die hoofstuk 4 ondersoek ek die effek van beweiding deur beeste (teenwoordigheid teenoor afwesigheid, sowel as beweidingsintensiteit) op die plantspesiesrykheid en samestelling van gebrande en ongebrande Afrikaberg grasvelde wat minder gereeld gebrand word. Die minste plant spesies is aangeteken in ongebrande, onbeweide grasveld in die NR. Brande en beweiding het albei 'n effek op plantspesiesamestelling gehad wat gepaard gegaan het met 'n toename in plantspesiesrykheid. Plantgemeenskappe in grasvelde wat onlangs (<12 maande voor die steekproef geneem is) gebrand is, het slegs van die in ongebrande grasvelde verskil wanneer nie een van die twee areas bewei is nie. Op 'n soortgelyke trant het die teenwoordigheid van beeste (EN teenoor NR) slegs n effek gehad in ongebrande grasvelde. Ongebrande plantgemeenskappe in die EN wat deur beeste bewei is, was baie soos die in die NR wat deur swartwildebeeste bewei is. Dit dui daarop dat beeste die effek van inheemse wildsoorte tot 'n mate naboots. Des nieteenstaande die bogenoemde, het swaar-beweide grasvelde minder plantspesies gehad as grasvelde wat slegs matig bewei is. Ek stel voor dat brande en beweiding deel moet vorm van die bestuur van grasvelde in EN’e, want hierdie natuurlike versteuringe dra by tot 'n diverse, dinamiese grasveldekosisteem. Tog moet bestuurders versigtig wees wanneer hulle die plaaslike gemeenskap se beeste in EN’e toelaat, want swaar beweiding kan bewaringsinisiatiewe in die wiele ry. In hoofstuk 5 het ek die klem na springkane verskuif, en die effek van jaarlikse brande, beweiding deur beeste (teenwoordigheid teenoor afwesigheid) en tydsverloop sedert laaste brand op hierdie sensitiewe insekte in Afrikaberg grasvelde ondersoek. Alhoewel springkaangemeenskappe baat gevind het by versteuringe, het hulle nie beduidend gereageer op enige van die indiwiduele versteuringe nie. Die digste sprinkaan bevolking met die hoogste spesies diversiteit is aangeteken in brandbane in die EN wat liggies deur beeste bewei is. Darenteen is die laagste bevolking en spesies diversiteit aangeteken in grasvelde in die NR wat groot hoeveelhede dooie plantmateriaal bevat wat aandui dat hierdie grasvelde nie onlangs gebrand het nie. Springkaangemeenskappe in gebrande grasvelde het slegs van ongebrande grasvelde verskil wanneer nie een van die twee bewei is nie. Die sleutelkombinasie van versteuringe wat die rykheid en samestelling van springkaangemeenskape bepaal het, was 'n hoë brandfrekwensie (soos in brandbane) en beweiding deur beeste. Wanneer een van hierdie versteuringe afwesig was, was springkaangemeenskappe tussen verskillende areas dieselfde. Alhoewel springkaangemeenskappe daarby baat gevind het wanneer brandbane elke jaar gebrand en deur beeste bewei is, kan ek nie hierdie bestuurspraktyk vir die res van die EN aanbeveel nie. Grasvelde in die res van die EN behoort eerder minder gereeld (elke 2-4 jaar) gebrand en met 'n rotasiestelsel bewei word. Sodoende sal brandbane voorsien in die behoeftes van springkane, en die res van die EN in die behoeftes van sensitiewe taksa wat onversteurde habitat benodig om te floreer. My slotsom is dat versteuringe nodig is om die volle diversiteit van plante en springkane en die dinamika binne-in grasvelde te bewaar. Tog verdwyn daar plantspesies uit areas met 'n hoë versteuringsintensiteit of frekwensie en klein, geïsoleerde kolle natuurlike plantegroei in die EN. Daarom beveel ek aan dat natuurlike versteuringe (brande en beweiding) matig toegepas moet word in wye, aaneengeskakelde gange in die EN. Hierdie benadering tot natuurbewaring kan biodiversiteit tussen bosbouplantasies beveilig teen verdere verlies.

Doutoramento em Ciências e Engenharia do Ambiente
O presente trabalho é constituído por uma revisão bibliográfica e uma série de estudos científicos experimentais visando a melhoria do conhecimento da ecologia de germinação e resposta ao estresse hídrico em arbustos autóctones. Estes estudos pretendem contribuir para o conhecimento da ecologia de espécies autóctones na fase de germinação e estabelecimento inicial das plântulas através da comparação da resposta de sementes de diferentes proveniências. Os primeiros três capítulos focam-se no estudo dos efeitos de pré-tratamentos na germinação de espécies arbustivas com diferentes estratégias de regeneração e dormência da semente, enquanto os dois últimos capítulos incidem sobre o efeito de diferentes condições de disponibilidade de água na germinação e estabelecimento inicial de plântulas com potencial para restauro ecológico. Este estudo procura abordar aspetos importantes para o estudo e validação do uso de espécies alvo para o restauro ecológico em zonas Mediterrânicas, contribuindo deste modo para a melhoria do conhecimento da ecologia destas espécies. A germinação foi estimulada por pré-tratamentos de calor em quatro das cinco espécies cuja época de dispersão da semente na primavera-verão. As sementes provenientes do sul tenderam a germinar melhor, sendo ao mesmo tempo mais pequenas. Por outro lado, a germinação em espécies cuja dispersão da semente ocorre no outono-inverno variou de acordo com o tipo de dormência da semente. A germinação foi em geral favorecida pela estratificação a frio na espécie apresentando dormência fisiológica mas foi negativamente afetada na espécie que não apresenta dormência. Esta resposta não foi, no entanto, inequívoca pois foi dependente da proveniência estudada. O efeito da estratificação a frio teve em geral um efeito mais positivo ou menos negativo nas sementes provenientes do norte em comparação com as outras proveniências. O stress-osmótico teve um efeito negativo consistente na germinação de Pistacia lentiscus e outro fator que também afetou significativamente a germinação foi a variabilidade intra-populacional entre plantas mãe. As duas proveniências apresentaram diferentes graus de associação entre a germinação e/ou caracteres morfológicos da semente ou planta mãe. Quanto à fase de estabelecimento inicial, o baixo conteúdo em água teve um claro efeito negativo no investimento relativo em biomassa acima do solo e um efeito positivo no investimento relativo em biomassa abaixo do solo em Arbutus unedo. Esta resposta não foi inequívoca, uma vez que as proveniências diferiram em adaptações morfológicas ao baixo conteúdo em água. As plântulas da proveniência mais húmida revelaram uma taxa de crescimento relativo superior mas, ao mesmo tempo, uma fraca adaptação às condições de baixa disponibilidade de água, quando comparadas com as outras proveniências. Ao contrário, as plântulas da proveniência seca com verão mais quente apresentaram semelhantes diâmetro do colo da raiz, peso seco de folhas e performance fisiológica sob os dois regimes de irrigação. Os resultados obtidos revelam diferenças significativas na germinação e desenvolvimento inicial entre as distintas proveniências que poderão estar relacionadas com adaptações ao clima dos locais de origem. Os desenvolvimentos futuros do estudo dos mecanismos subjacentes às adaptações observadas poderiam contribuir para a melhoria das previsões de sucesso do estabelecimento inicial em diferentes populações de plantas, permitindo aumentar a confiança e efetividade de custos nas decisões relacionadas com ações de restauro ecológico em cenários de alterações climáticas.
The present work is composed by a conceptual scheme based on a literature review plus a series of experimental scientific studies. These studies aim at improving the knowledge base of germination ecology and response to water stress in autochthonous shrub species through the comparison of different provenances. The first three chapters focus on the study of the role of pretreatments in the germination of shrub species with different regeneration and seed dormancy strategies while the two last chapters focus on the study of the effect of contrasting water availabilities in the germination and early establishment of species with restoration potential. This study intends to address important aspects for the study and validation of the use of target species for ecological restoration actions in Mediterranean areas, thereby contributing to the improvement of the knowledge of the ecology of these species. Germination in four out of the five species whose seeds are dispersed in spring-summer was enhanced by heat pre-treatments. The seeds from southern provenances tended to germinate better, being at the same time smaller. On the other hand, germination response in autumn-winter seed dispersed species varied according to the type of seed dormancy. Germination was, in general, enhanced by cold stratification in the species presenting physiological dormancy but negatively affected in the species with no dormancy. This response was, however, not unequivocal and depended on the role of provenance. The effect of cold stratification had, in general a more positive or less negative effect in the germination of the northern seeds when comparing to the other provenances. Osmotic-stress had a consistent negative effect on germination of Pistacia lentiscus and another factor that significantly influenced germination was intrapopulation variability between mother-plant. The two provenances presented distinct degrees of association between germination and seed or mother plant morphological traits. As to the early establishment phase, low water content had a clear negative effect on Arbutus unedo seedlings’ aboveground biomass and a positive effect on belowground biomass relative investment. This response was not unequivocal, since provenances differed in morphological adaptations to low water content. Seedlings from the wettest provenance revealed a higher relative growth rate under high water content but a poor adaptation to limited water availability when compared to the other two provenances. By contrast, seedlings from the dry provenance with the hottest summer had similar root collar diameter, leaf dry weight and physiological performance under both watering regimes. The present results revealed significant differences in germination and early development response among different provenances that could be related with adaptations to the climate of the seed source. Further studies on the mechanisms underlying the observed adaptations could contribute to the improvement of early establishment success predictions in different plant populations, allowing more reliable and cost-effective management ecological restoration-related decisions under climate change scenarios.

The aim of this thesis is to examine the role of anthropogenic disturbance in the creation of a socio-ecological landscape. Three key questions were answered: what impact has past anthropogenic disturbance had on present vegetative characteristics; what value did this disturbed landscape have to local people; how did the local peoples' worldviews and eco-cosmologies influence how they perceived, valued and managed their landscape? Research was based in a rural, predominantly amaXhosa village in the Kat River valley, Eastern Cape, South Africa. Four major landscape components characterised the environment, namely dense forests, former grazing lands, abandoned fields and old settlements. A combination of qualitative and quantitative methods were adopted, consisting of a vegetation and soil survey and social science methods including semistructured interviews, focus groups, transect walks and participant observation. The key finding was that anthropogenic disturbance is necessary to enhance the potential of the area to support human habitation. However, anthropogenic disturbance can have positive or negative effects for both local people and the environment. An intermediate level of disturbance is a key factor leading to a resilient socio-ecological system. Various anthropogenic disturbances have had significant affects on vegetation characteristics in terms of species richness, and a change in vegetation composition and species heights. Of the three anthropogenic landscapes examined, former grazing lands were the least intensively disturbed. They were also more species rich and structurally diverse than areas that were under agricultural production or used as a settlement. Dense forests, although the least desirable landscape to local people were, together with sacred pools, sources of ecosystem renewal and played a critical role during times of disturbance. Anthropogenic disturbance has added value to local people's livelihoods; this disturbance has resulted in a patchy landscape that supplies a wide variety of resources to local people. Local people are dependent on their environment for their basic needs; this has encouraged sustainable management practices. The local AmaXhosa still retain many elements of their traditional worldview, which has many characteristics of Animism. The environment is an integral component of their belief system and certain sites and species have significant cultural value and are protected.

Along the 80-km gravel reach between Hope and Mission, Fraser River has a wandering morphology that includes secondary channels, gravel bars, and islands that together support a diverse and productive ecosystem. The wandering morphology is produced, in part, by annual sediment deposition within the reach. Sediment deposition, however, is perceived as a threat to flood security and gravel mining is proposed as a profitable solution to flood risk. This thesis presents a hierarchical habitat classification for the gravel reach, which provided a spatial framework to, first, examine habitat associations of benthic invertebrates and fish and, second, evaluate the physical and ecological responses to habitat disturbance by gravel mining. At the highest level of the classification, the river is divided into 5 sub-reaches (104 m scale) that vary in morphological expression and sediment gradational tendency. The intermediate level specifies gravel bar units (103 m scale) each consisting of a riffle, gravel bar, and adjacent pool. Nested within gravel bars are physically distinct habitat units, which represent the finest level of the classification (101- 102 m scale). Results demonstrated that the assemblages of invertebrates and fish associated with habitats are moderately distinct and differentiated along a hydraulic gradient corresponding to velocity. However, the congruence between habitat structure and the structure of aquatic communities was weakened by large spatial and temporal variability in the distribution and abundances of many species. This variability was predicted to afford the invertebrate and fish community resilience to physical disturbance. The second component of this study examined the effect of disturbance by gravel mining from an exposed bar in 2000. Physical changes immediately after mining were substantial as the pre existing cobble surface was replaced by loose gravel and sand. Two subsequent freshets transformed the site into a topographically complex area with similar substrate texture as before mining, although sediment replenishment to the site was negligible. Habitat availability at flows <3000 m3 s-1 increased and habitat quality was comparable with reference sites. A third above-average freshet replenished 31% of the removal volume and restored average bar surface elevation to within 9 cm of the pre-scalped surface. Physical changes elicited a significant reduction in invertebrate density immediately after mining, however, the impact lasted less than one freshet cycle. No change in fish density as a result of mining was found, although statistical power to detect an impact was low and fish sampling was carried out at flows <5700 m3 s-1. These results support the expectation that the invertebrate and fish community in the gravel reach has resilience to disturbance from a single gravel removal provided that site recovery by way of sediment transport and replenishment occurs.
Arts, Faculty of
Geography, Department of
Graduate

Temperate forest ecosystems are subject to various disturbances including insect agents, drought and fire, which can have profound effects on the structure of the ecosystem for many years after the event. Impacts of disturbance can vary widely, therefore an understanding of the legacies of an event are critical in the interpretation of contemporary forest patterns and those of the near future. The primary objective of this dissertation was to investigate the ecological legacies of drought, beetle outbreak and ensuing wildfire in two different ecosystems. A secondary objective of my research, data development, was motivated by a lack of available data which precluded ecological investigation of each disturbance.

I studied the effects of drought on deciduous and coniferous forest along a forest-shrubland ecotone in the southern portion of the Wyoming Basin Ecoregion. The results show that forests in the region have experienced high levels of cumulative drought related mortality over the last decade. Negative trends were not consistent across forest type or distributed randomly across the study area. The patterns of long-term trends highlight areas of forest that are resistant, persistent or vulnerable to severe drought.

In the second thread of my dissertation, I used multiple lines of evidence to retrospectively characterize a landscape scale mountain pine beetle disturbance from the 1970s in Glacier National Park. The lack of spatially explicit data on this disturbance was a major data gap since wildfire had removed some of the evidence from the landscape. I used this information to assess the influence of beetle severity on the burn severity of subsequent wildfires in the decades after the outbreak. Although many factors contribute to burn severity, my results indicate that beetle severity can positively influence burn severity of wildfire. This is likely due to the change in forest structure in the decades after the outbreak and not as a direct result of tree mortality from the outbreak. The long-term perspective of this study suggests that ecological legacies of high severity disturbance may continue to influence subsequent disturbance for many years after the initial event. This work also provides insight on future disturbance interactions associated with the recent mountain pine beetle outbreak that has impacted tens of millions of hectares in western North America over the last two decades.

The main cause of environmental disturbance in the Fennoscandian boreal forests today is forestry. Natural disturbances are important to maintain diversity, but anthropogenic disturbance, such as forestry, differs in many ways from the natural ones. Forestry is a big industry in Sweden and only a small remnant of old-growth forest is left. Several studies have shown an initial decrease in e.g. dragonfly diversity a few years after logging, followed by an increase up to numbers comparable with the original species number. In this study I examined whether the new, quite diverse, species composition is similar to the one present before the logging or if some species are disappearing and are replaced with other, maybe opportunistic species. Other factors such as ongoing changes in climate also will be considered. A resampling of 34 (and an additional 4) lakes that also were sampled in 1996-97 was done during summer of 2006. Analyses of data from the two sampling occasions were done. I found that even if the diversity is just temporarily affected (or not affected at all), it is not always the same species involved. This means that the diversity in a single lake can appear to be high, but the total diversity in Sweden, or Scandinavia, is declining. I also found some interesting new species for the area, such as Nehalennia speciosa, Sympecma fusca and Aeshna mixta.

Disturbance from engineering works is an increasing problem in terrestrial and marine ecosystems throughout the world. Many reported declines in population size, breeding success and body condition have been diagnosed as the result of anthropogenic disturbance, however little is known about the effect of long-term disturbance from large-scale engineering works. Understanding the mechanisms by which animals respond to anthropogenic activities is fundamental to explaining interactions, and resolving potential conflicts between humans and wildlife. This thesis focuses on the factors affecting the habitat use and foraging decisions in wintering shorebirds and wildfowl. The first half of this thesis considers the direct and indirect impacts on waterbirds of a major engineering project in central Scotland; construction of the new Clackmannanshire Bridge at Kincardine-on-Forth. For individual bird species in close proximity to the bridge site, round-the-clock construction work had consequences ranging from neutral to considerably negative. Cormorant Phalacrocorax carbo declined in the area, probably as a result of the disturbance of an important low tide roost. Redshank Tringa totanus, previously abundant in the prey-rich areas adjacent to the construction site, were displaced into poorer areas for most of the construction period; where they may also have suffered from increased interference competition and elevated risk from raptorial predators. Some positive effects of industrial development were also revealed; radio-transmitters combined with tilt-switch posture sensors indicate that Redshank were able to capitalise on the improved nocturnal visibility in areas around Grangemouth docks to assist with foraging and predator detection. Evidence is presented that birds switched foraging strategy (from sight to touch feeding) depending on ambient light levels; whereby artificial light was used in a similar manner to moonlight to assist with prey detection. Redshank also avoided riverine areas at night that were used frequently by day, probably in response to an elevated threat from nocturnal predators. As the predator landscape changes from day into night, birds adopt different strategies to minimise the risk from nocturnal predators. It is clearly important, therefore, that information on nocturnal distributions is available to inform decisions on site management, especially where anthropogenic activity continues throughout the diel cycle. Behavioural decisions were shown to vary widely within a species depending on individual state, metabolic demands and previous exposure to human disturbance. Prey resources were shown to change dramatically over the course of a winter. In response to this decline, the home range of Redshank contracted over a winter season. Similarly, animals responded less and took greater risks in response to experimental disturbance events later in the winter than earlier in the winter, and on days when the temperature was lower. This effect was strongest for individuals occupying heavily disturbed areas, which were possibly already compensating for lost feeding time and a negative energy balance. The results were consistent with the hypothesis that those individuals that respond most obviously to human disturbance were those least likely to suffer fitness consequences. This is the opposite from what is commonly assumed when behaviour is used as an index of disturbance impacts, most notably in the use of flush distance in the design of wildlife buffer zones. In conclusion, this study demonstrated various negative impacts of disturbance, including local displacement, due to construction activity on overwintering waterbirds. It also revealed two key, but poorly understood, phenomena relating to mechanisms for coping with anthropogenic disturbance: routine utilisation of artificial light to extend night-time feeding opportunities amongst Redshank and an adaptive flexibility in escape responses across a range of species under varying conditions of risk.

This qualitative study addressed an intersection in which the application of the medical model to educational disabilities and its implications for educational labeling of students interacted with the professional enculturation of clinical social workers. Employed as clinicians in programs administered in a large south eastern school district in the United States, five social workers served elementary students labeled Emotionally/Behaviorally Disabled (EBD) through federally authorized provisions for special education related services. This study used grounded theory methods to discover and analyze the social workers' underlying assumptions, values, and patterns of practice with regard to ethical and guild issues, roles and responsibilities, and bases of knowledge. The study found that participants interpreted and applied a knowledge base grounded in the normative aspects of social work. Their preparation made it possible to compete for licensure (LCSW) and assume roles as professional helpers but did not provide all the tools they needed to carry out their work as clinicians with students in EBD programs. Secondly, it found that ambiguities regarding ethics, guild issues, and roles emerged with regard to acting as helping professionals in an integrated professional setting. Each practitioner exercised certain latitude to respond as needed to challenges which varied from site to site. Finally, the findings reflected how the clinicians have situated themselves in the face of the demands of documentation procedures and of participation in meetings endemic to special education.

Peat mosses (Sphagnum) are widely distributed and are a major component of mire vegetation and peat throughout the boreal and temperate regions. Most boreal Sphagnum species regularly produce sporophytes, but the ecological role of the spore has been questioned. This study shows that the spores can form a spore bank and have the ability to germinate and contribute to moss establishment whenever suitable conditions occur. The results suggest that spore production is important for explaining the wide distribution and omnipresence of Sphagnum in nutrient-poor wetlands. The results further imply that initial recruitment from spores predominates in Sphagnum after disturbance or formation of suitable habitats.

A series of experiments showed that addition of phosphorus-containing substrates, such as fresh plant litter or moose dung, resulted in spore establishment on bare, moist peat. A field experiment indicated establishment rates of about 1% of sown, germinable spores on peat with added substrates. Plant litter on moist soil, without a closed cover of bryophytes, is an important safe site for the establishment of Sphagnum spores. The results fit the observed pattern of colonisation by Sphagnum beneath Eriophorum vaginatum tussocks in mires severely disturbed by peat extraction. Successful long-distance dispersal was indicated by the occurrence of several regionally new or rare Sphagnum species in disturbed mires.

Spore number per sporophyte ranged among Sphagnum species from 18 500 to 240 000, with a trade-off between spore number and spore size. Annual spore production was estimated at 15 million spores per square metre on two investigated mires. Sporophyte production showed a large interannual variation. Sporophyte production was positively related to the amount of precipitation the preceding summer. This was probably because a high water level promoted gametangium formation. Spore dispersal occurred in July and August. The earlier timing of spore dispersal in the more drought-sensitive, hollow-inhabiting sphagna should reduce the risk of sporophytes drying out prematurely during summer droughts.

Spores kept refrigerated up to 13 years retained high germinability. A field experiment showed that Sphagnum can form a persistent spore bank, with a potential longevity of several decades.

Tatera brantsii was numerically dominant in the small mammal community on the plateaux of the rehabilitated ash disposal sites of ESKOM's Hendrina Power Station in 1998 and 1999 (Vermaak 2000). The species seemed well adapted to exploit this environment and, through biopedturbation, had altered the topsoil structure and chemistry. The consequences of this and other activities also affected the rehabilitated plant community of the PFA-dam habitat. Burrowing appeared limited to just under the topsoil layer, and seemed more extensive than burrows of this species in natural ecosystems. The burrow system architecture was mapped and quantified, and localised increases in nitrates, phosphorous and organic carbon in immediately associated substrate were noted. However, this substrate enrichment was transient, and disappeared following the abandonment, and subsequent collapse of burrow systems, when gerbil colonies migrated away from the area. The mixing of soil horizons also resulted in a more homogeneous substrate, which was more friable, and thus drier. The high pH and salinity of the topsoil layer in areas undisturbed by gerbil burrowing, and concentrations of particular elements associated with either the topsoil covering or the ash, were reduced as a consequence of substrate mixing in disturbed areas. Gerbil impacts on the substrate of this habitat seemed to promote pedogenesis, eliminating the sharp distinction between the topsoil covering and the ash below, but the re-exposed ash of the burrow mounds would become subject to erosion, and reduce the effectiveness of the rehabilitation effort. Gerbil activities increased the number of plant species, especially ruderal forbs, comprising the plant community of the PFA-dam habitat, but plant community diversity was not significantly increased. However, numerical dominance by few tussock grass species was diminished , possibly reflecting burial under mounds of excavated substrate. The biomass and cover of some grass species were reduced in areas of gerbil impacts, and plant lifecycles appeared to be completed sooner in areas affected by gerbil activities. These effects may be as a result of the drier substrate produced following the collapse of the extensive network of abandoned burrows. The succession of this plant community towards an underutilised grassland state, the expected outcome of the rehabilitation effort, was minimally affected by gerbil activities. The effects of T.brantsii activities in this PFA-dam habitat were not as distinct as the effects noted by other authors studying fossorial rodent impacts in less disturbed habitats. This could be because further disturbances in this habitat would merely add to the currently disturbed state, whereas disturbance in more natural habitats, would show more of a change from the initial state.
Thesis (M.Sc. (Zoology))--North-West University, Potchefstroom Campus, 2007.

Human activities and expansion have created novel disturbance patterns across Earth’s landscapes. Disturbance is an ecological interruption after which ecosystem recovery or transition into a new state can occur, affecting biodiversity, ecosystem functioning, and theavailability of ecosystem services. Fire and herbivory are two of the most widespread forces of disturbance which shape ecosystems globally. In temperate forest ecosystems, fire affects forest composition, which influences the diversity and resilience of ecosystems (chapters 1 and 2) and forest canopy structure, which is important to snowpack accumulation and the availability of water resources (chapters 3 and 4). In chapter one, the effects of conifer competition, which occurs under fire suppression, and ungulate herbivory on aspen regeneration and recruitmentwere examined. It was found that conifer competition, and ungulate herbivory both drastically reduced successful aspen regeneration and recruitment and had a larger effect than climatic or topographical variables. In chapter two, this understanding was used to investigate mechanicaland fire interventions by the National Forest Service in a mixed aspen conifer forest experiencing fire suppression and heavy ungulate herbivory. Untreated forests failed to recruit aspen suckers successfully due to conifer competition and ungulate browsing. Fire treatments were successful in restoring aspen habitat, but mechanical treatments failed due to heavy ungulate use, despitethe treatment eliciting high sucker densities. In chapter three, fire severity was found to have important implications for snowpack accumulation and snow-water content in alpine forests. High burn severity, which is projected to become more common under future climaticconditions, led to deeper and denser snowpack possibly buffering the effects of water loss in a warmer climate. In chapter four, the interaction between topography and vegetation in burned forest conditions was evaluated. It was found that topographical aspect likely mediates the effect of vegetation on snowpack and may have an opposite effect on snow accumulation and melt on north vs south facing aspects. A synthesis of studies from different regions further supports the idea that this relationship between fire and snow is heavily dependent on latitude, elevation, and slope angle. Together, these findings demonstrate that the resilience and persistence of aspenforest ecosystems in changing disturbance regimes depend on complex interactions among disturbance type, disturbance severity, landscape position, and hydrology. These interactions should be integrated into management strategies to protect ecosystems and ensure ecosystemservices for growing human populations in the western United States.

Climate change is increasing the frequency and extent of high-severity disturbance, with potential to alter vegetation community composition and structure in environments sensitive to tipping points between alternative states. Shrub species display a range of characteristics that promote resistance and resilience to disturbance, and which yield differential post-disturbance outcomes. We investigated differences in shrub patch size and stem density in response to variations in fire severity, vegetation community, and post-disturbance reproductive strategies in Sky Island forested ecosystems in the southwestern United States. Patterns in shrub structure reflect the effects of fire severity as well as differences among species with alternate post-fire reproductive strategies. Increased fire severity correlates with larger patch sizes and greater stem densities; these patterns are observed across multiple fire events, indicating that disturbance legacies can persist for decades. High severity fire produces the largest shrub patches, and variance in shrub patch size increases with severity. High severity fire is likely to promote expansion of shrub species on the landscape, with implications for future community structure. Resprouting species have the greatest variability in patch structure, while seeding species show a strong response to disturbance: resprouting species dominateatlowdisturbanceseverities,andobligateseedersdominatehighseverityareas. Differential post-fire reproductive strategies are likely to generate distinct patterns of vegetation distribution following disturbance, with implications for community composition at various scales. Shrub species demonstrate flexible responses to wildfire disturbance severity that are reflected in shrub patch dynamics at small and intermediate scales.

A great deal of research has demonstrated that Non-Timber Forest Products (NTFPs) play a crucial role in the livelihoods of the rural poor, and are particularly important to the most marginalised people throughout the developing world. However, these livelihood benefits are not without cost to the natural resource base that rural communities depend so heavily upon. The continued dependence on NTFPs as a major livelihood source must be contingent upon the minimisation of the level of disturbance created through this dependency. This study assesses the level of disturbance created through natural resource harvesting in eight study sites around South Africa, and applies a predictive conceptual model created by Shankaar et al. (2004b) to try and ascertain under what conditions the level of disturbance created through natural resource harvesting will be high. It assesses the three key factors that Shankaar et al. (2004b) identified (level of Local Ecological Knowledge (LEK), level of dependency and access to markets) in relation to the level of disturbance found at each of the study sites. It was found that there was a statistically significant relationship between the level of dependency and the level of disturbance, but there was no statistically significant relationship between either access to markets or the level of LEK and disturbance. Regulation of land use is a key issue, with weak local institutions in communal areas making effective resource management difficult. The significance of these findings is discussed, and priorities for future research are identified. This study adds to the body of knowledge related to NTFP harvesting and critically analyses the conflicts between the livelihood gains and the level of disturbance created through NTFP harvesting in an attempt to ascertain how livelihoods can be safeguarded. And in the longer-term, so that management strategies can be identified where resource extraction is not at the cost of undermining the very livelihoods that depend upon the natural resource base.

This study analyzes data acquired in French Polynesia in the Pacific and The Bahamas (Atlantic), both oceans affected by recent, well documented and sequential disturbances. For the purposes of this study, a disturbance is defined as a perturbation of environmental, physical or biological conditions that causes a distinct change in the ecosystem. After several decades of coral bleaching events, biological change, and anthropogenic impacts, rapid assessments of the coral community were accomplished by collecting photo-transects across the reefs to extract size structure of the corals, percent live tissue cover and perform a faunal evaluation. Cluster analyses and spatial autocorrelation tests were done to examine the community structure and dynamics at both locations. All multivariate analyses pointed to a disturbed ecosystem and the lack of spatial correlation indicated the impact of a local disturbance over that of a regional event. In assessing the spatial coral community structure, different responses to large versus small scales of disturbance were found. This emphasizes the importance of tailoring management of coral reefs to specific impacts. These two distinct regions were shown to have correlated spatial response patterns to sequential disturbances, supporting the idea of community pattern signatures for different scales of disturbance and the need for an adjustment in management protocols.

No contexto da aplicação da Directiva-Quadro da Água, a ictiofauna é considerada como um relevante elemento biológico. Pela primeira vez foi desenvolvido um sistema de classificação para avaliação do estado ecológico dos ecossistemas aquáticos em Portugal, apesar dos constrangimentos impostos pela variedade e ubiquidade das pressões humanas e pelas peculiaridades dos agrupamentos piscícolas. Foram definidos grupos piscícolas para posterior estabelecimento de uma tipologia nacional de rios, foram seleccionadas métricas responsivas ás pressões humanas e foi desenvolvido um índice biótico piscícola. Os resultados evidenciam a necessidade de diferenciar correctamente a resposta da ictiofauna ás perturbações naturais e antropogénicas, considerando quer a variabilidade espacial, quer temporal dos agrupamentos piscícolas, típica dos cursos de tipo Mediterrânico. O uso de solo tem fortes consequências na degradação dos rios e na integridade da ictiofauna, sendo importante que a avaliação dos impactos considere a interacção entre diferentes pressões. A compreensão da influência relativa dos factores ambientais e humanos na ocorrência e abundância das espécies não nativas permitiu evidenciar o potencial papel das mesmas como indicadores biológicos. Esta tese contribuiu para o desenvolvimento e aperfeiçoamento de ferramentas de avaliação ecológica em Portugal, permitindo igualmente um aumento do conhecimento sobre a ecologia das comunidades piscícolas e funcionamento dos ecossistemas em rios de tipo Mediterrânico, com vista à sua conservação e reabilitação; ABSTRACT:Under the implementation of the Water Framework Directive fish fauna is considered a useful biological element. For the first time a Portuguese classification system to assess the ecological status of aquatic ecosystems was established, despite several constrains posed by the variety and ubiquity of human pressures and by the peculiarities of the fish assemblages. It was possible to define fish-based geographical groups for the subsequent establishment of a national river typology, to select metrics responsive to human pressures and to develop a fish biotic index. Results underline the needs for accurately differentiate between fish responses to natural and anthropogenic disturbances by accounting for the spatial and temporal variability of fish assemblages. Land use may lead to strong impacts on stream degradation and fish assemblages integrity and this evaluation must consider the interaction of different pressures. Understanding the influence of environmental and human factors in the occurrence and abundance of non-native species enlightened their role as a reliable tool in the ecological assessment of rivers. This thesis represents an important contribution to improve the ecological assessment tools in Portugal, while increasing knowledge on the ecology of fish communities and ecosystem functioning in Mediterranean climate rivers in view of their urgent conservation.

Forest disturbance occurs on a wide gradient of selectiveness and creates new growth opportunities for adapted species. Across the spectrum of disturbance, anthropogenic disturbance influences community assembly in the Midwest more than other mechanisms but is its role in shaping and maintaining ecosystems is inadequately considered in most discussions on the historic range of variability (HRV). Forest resiliency is threatened by unprecedented agents of ecosystem change such as invasive species and reduced regeneration potential of native species. Historic anthropogenic disturbance largely resulted in forest conditions which commonly contained high value attributes like heterogeneity across habitat types and landscape diversity, yet also produced forests of undesirable traits due to high grading for timber and overgrazing by domesticated stock. In order to maintain historical representative forests and improve the degraded forests, active forest management is necessary to continue historic disturbance patterns and combat new threats. Forest transition theory is used here to describe the impacts of human settlement and development activities on forest ecosystems across the Middle Mississippi River Valley. To date, researchers have identified the need for information related to changes of forest attributes such as species composition and stand structure, improved descriptions of short- and medium-term dynamics within the context of the long-term transition, and the integration of biophysical drivers of forest change through time. In Midwestern U.S.A., forest dynamics were influenced by frequent, low intensity disturbance events that mediate forest composition and stand structure by selecting for disturbance regimes that create oak woodland and interspersed prairies and meadows. The onset of Euro-American settlement was accompanied by detailed land-use records with information related to forest attributes, agricultural activities, and parcel ownership patterns. We aggregated multiple sources of historic forest conditions into a geodatabase in order to document changes over the past 200 years in Elsah Township, Illinois, where the pre-settlement (1820) forest, once dominated by oak and hickory species, has largely shifted to a maple dominated system with a declining oak-hickory component, heavily influenced by an invasive shrub species, bush honeysuckle. Using on ordinary kriging interpolation, forest density was estimated at 8.7 stems per acre on average with a mean basal area of 14.6 square feet per acre prior to settlement. Conservation practices of the early 1900s, including fire suppression and erosion control resulted in changes to forest structure with density increases to 127 trees per acre with a basal area of 175.8 square feet per acre. The high degree of topographic variability near the Mississippi River influenced forest cover changes as slopes with low angles were the first to be converted from forest cover to other land uses (circa 1850). Forest re-initiation occurred in areas with steeper slope due to a lack of human activities. Forest cover declined to the lowest point in 1927 and has been rebounding steadily throughout this century. Of the original 15,252 forested acres, 11.6% remained covered throughout the past 200 years and coincided with slopes with an average of 39.1 degrees. These data can provide a spatially explicit and historically accurate tool to guide land management decisions including restoration treatment, disturbance regime management, and land use preservation activities in similarly heterogeneous environments. Forest communities along the bluffs of the Mississippi River differ in species composition and stand structure associated with specific topographic positions of floodplain, transition talus slope, bluff top, and upland. In order to assess current stand characteristics and ecosystem trajectory, we measured all woody stems in 316 fixed radius plots (79 plots per topographic position) with a plot area of 25 m2. Alpha (defined as within system diversity) and Beta (defined as between system diversity) diversity and diameter distributions were determined for seedling, shrub layer, and overstory stems. Stem density increased from 21.4 stems ha-1 in 1820 to 613 stems ha-1 in 1936 followed by reduction to 314 stems ha-1 in 2017. Average stand diameter decreased from 40.9 cm in 1820 to 25.3 cm in 2017 (for upland stems greater than 7.5 cm) while basal area increased from 3.3 m2 ha-1 in 1820 to 40.4 m2 ha-1 in 2017. Alpha diversity was highest in the upland overstory and in the river island shrub layer. Beta diversity in the overstory was highest (0.67) between the bluff and the upland while lowest (0.08) between the bluff and the river island. Importantly, mesophytic species are no longer restricted to watercourses and valleys as reported in historical accounts and confirmed by the spatial analysis of original witness tree records. Currently, bush honeysuckle, an invasive species, dominates the shrub layer on most non-hydric sites of the talus slope, upland, and particularly across the bluff top where it is an indicator. Across all forest sites in the study, we found evidence of a community shift to less diversity and more mesophytic species over the past 80 years. Hill prairie vegetation on the limestone bluffs of the central Mississippi River Valley represents a significant portion of the remaining prairie, savanna, and woodland systems of the Midwest and should be appropriately managed with prescribed fire and woody stem reduction efforts. We examined the structure, composition, and temporal community patterns of the forest-prairie gradient by employing hierarchical cluster analysis and non-metric multi-dimensional scaling in combination with indicator species analysis and dendrochronological methods. Results suggest that four general community types exist across the forest-prairie gradient: Group 1 consists of the woodland community structure with significant indicator values for the density of Juniperus virginiana (indicator value 58.4, p = 0.0002), Carya glabra (45, 0.0022), Quercus stellata (23.7, 0.0424), and Lonicera maackii (74.2, 0.0002) and a high basal area (BA) of J. virgniana (21.4, 0.0276) and L. maackii (47.9, 0.0054). The first year of L. maackii presence was 1964 with the primary wave of invasion beginning around 1990. Group 2 contains bare soil coverage in the subplot (40.4, 0.0002) as the one indicator at a significant level. The species with the highest BA in Group 2 include Acer saccharum (9.08 m2 ha-1), Q. velutina (5.89 m2 ha-1), and Q. muehlenbergii (5.32 m2 ha-1). Group 3 typifies the hill prairie community with the sole indicator of grass coverage in the subplots (39.7, 0.0196). Group 4 represents the stage of forest development following the cessation of disturbance events and the trajectory advancing towards a mesophytic forest and contains 14 significant indicators.

Future climate projections suggest an increase in average temperature as well as a decrease in average winter rainfall across the south-west Australian floristic region (SWAFR). These adverse future climatic conditions will amplify the intensity and frequency of disturbance events such as drought and fire. Mediterranean forests within the SWAFR are prone to drought and fire disturbance and have acquired resilience through the selection of drought and fire tolerable species. However, shifts in the magnitude of these disturbance events could increase the recovery period required for recruitment, causing a shift in forest structure and decreasing the resilience of these ecosystems to future disturbances. In this study, we investigated above-ground biomass (AGB) accumulation of understorey plants at sites within the Northern Jarrah Forest (NJF) that have experienced different degrees of drought and fire intensity. We found that within a disturbance event, sites experiencing either more severe drought and fire intensities on average accumulated substantially more understorey AGB than sites subjected to both low drought and moderate fire intensities. This suggests that understorey species within the SWAFR gain a competitive advantage in high drought and fire severity conditions and are highly tolerant to drought. However, the increase in understorey AGB accumulation also suggests a shift in overall forest structure to more dense, compact, low-ground small stems, which is known to increase fire probability. An increase in fire probability shortens the time period between fire intervals and can detrimentally affect forest recovery, especially in drought conditions. Therefore, these changes may shift ecosystems within the SWAFR to a state of non-equilibrium and reduce resilience to future disturbance events.

xii, 64 p. : ill. (some col.), maps
Salmon habitat restoration is ongoing at a Nature Conservancy preserve on the Middle Fork John Day River in the Columbia River Basin in north-central Oregon. The site has a long history of disturbance including dredge mining upstream, channelization, grazing, logging, fire, and floods. Using historic aerial photos, habitat unit surveys, and cross sectional profiles, this thesis shows how the channel morphology, particularly habitat unit diversity, has changed since 1939, just before placer mining began. Results show that the dominant influence on present day channel morphology is channelization from the 1930's. Other changes including dredge mining in the late 1930's to early 1940's, cessation of cattle grazing in 1991, and a fire followed by a flood in the winter of 1996-1997, had less impact because of the straightened, stabilized channel morphology.
Committee in charge: Patricia McDowell, Chairperson; W. Andrew Marcus, Member
10000-01-01

The ecological restoration of Peninsula Shale Renosterveld is essential to redress its conservation target shortfall. The ecosystem is Critically Endangered and, along with all other renosterveld types in the Cape lowlands, declared 'totally irreplaceable'. Further to conserving all extant remnants, ecological restoration is required to play a critical part in securing biodiversity and to meeting conservation targets. Remnants of Peninsula Shale Renosterveld are situated either side of the Cape Town city bowl and, despite formal protection, areas of the ecosystem are degraded and require restoration intervention. The body of research addressing restoration in renosterveld is small, yet growing, and seed-based restoration efforts have achieved limited success. This study primarily set out to further understand community recovery through determining the effects of interventions, implemented to mimic ecological drivers, on several seed germination criteria. Thirty-two interventions (comprised of five, crossed factors: burning, tillage, herbicide-application, rodent-exclusion and seeding) were incorporated into a field experiment situated in an area that was most likely ploughed over a century ago and is currently dominated by alien, annual grasses. Additional aspects of the study included determining the physiological status of the seed from 31 harvested species (through viability and germinability tests) towards identifying key restoration species, and, assessing intervention costeffectiveness as a measure of intervention feasibility. The majority of the harvested species exhibited moderate to high levels of viability and germinability and occurred in the middle or upper key-restoration-species index range, indicating their potential for use in future restoration efforts. Seeding contributed considerably to overall community attributes, significantly increasing indigenous seedling density, species richness and canopy cover. Due to the dominance of alien, annual grasses, seeding alone was relatively ineffectual and recruitment was considerably improved when seeding was implemented in conjunction with one or more of the other interventions. These findings indicate that a lack of available seed is not the only barrier to community recovery and that competition exerted by the alien grass component plays a large role in inhibiting seedling recruitment of desirable species. Intervention effectiveness increased with the number of factors per intervention yet, fortuitously, the most effective interventions were not necessarily the most costly. Some interventions resulted in good performances and have the potential to restore self-perpetuating communities with a semblance of ecosystem composition, structure and function. There is a clear and promising way forward incorporating these findings into feasible, implementable, landscape-scale, ecological restoration strategies for Peninsula Shale Renosterveld and potentially other Critically Endangered and 'totally irreplaceable' renosterveld ecosystems.

As climate change continues, climactic extremes are predicted to become more frequent and intense, in some cases resulting in dramatic changes to ecosystems. The effects of climate change on ecosystems will be mediated, in part, by biotic interactions in those ecosystems. However, there is still considerable uncertainty about where and how such biotic interactions will be important in the context of ecosystem disturbance and climactic extremes. Here, I review the role of consumers in seagrass ecosystems and investigate the ecological impacts of an extreme climactic event (marine heat wave) and subsequent widespread seagrass die-off in Shark Bay, Western Australia. Specifically, I compare seagrass cover, shark catch rates, and encounter rates of air breathing fauna in multiple habitat types before and after the seagrass die-off to describe post-disturbance dynamics of the seagrass community, shifts in consumer abundances, and changes in risk-sensitive habitat use patterns by a variety of mesoconsumers at risk of predation from tiger sharks (Galeocerdo cuvier). Finally, I conducted a 16 month field experiment to assess whether xi loss of top predators, and predicted shifts in dugong foraging, could destabilize remaining seagrass. I found that the previously dominant temperate seagrass Amphibolis antarctica is stable, but not increasing. Conversely, an early-successional tropical seagrass, Halodule uninervis, is expanding. Following the die-off, the densities of several consumer species (cormorants, green turtles, sea snakes, and dugongs) declined, while others (Indo-Pacific bottlenose dolphins, loggerhead sea turtles, tiger sharks) remained stable. Stable tiger shark abundances following the seagrass die-off suggest that the seascape of fear remains intact in this system. However, several consumers (dolphins, cormorants) began to use dangerous but profitable seagrass banks more often following seagrass decline, suggesting a relaxation of anti-predator behavior. Experimental results suggest that a loss of tiger sharks would result in a behaviorally mediated trophic cascade (BMTC) in degraded seagrass beds, further destabilizing them and potentially resulting in a phase shift. My work shows that climactic extremes can have strong but variable impacts on ecosystems mediated in part by species identity, and that maintenance of top predator populations may by important to ecological resilience in the face of climate change.

In this thesis, I combined field, common-garden and greenhouse experiments to examine the ecological and evolutionary consequences of plant-herbivore interactions in the perennial herb Lythrum salicaria. More specifically I examined (1) whether resistance and tolerance to damage from herbivores vary with latitude and are positively related to the intensity of herbivory in natural populations, (2) whether effects of herbivory on plant fitness vary with latitude, (3) whether populations are locally adapted and whether herbivory influences the relative fitness of populations, and (4) whether the intensity and effects of insect herbivory on reproductive output vary locally along a disturbance gradient and are associated with differences in plant resistance. A common-garden and a greenhouse experiment demonstrated that plant resistance decreased whereas plant tolerance increased with latitude of origin among populations sampled along a latitudinal gradient in Sweden. Oviposition and feeding preference in the greenhouse and leaf damage in the common-garden experiment were negatively related to natural damage in the source populations. Experimental removal of insect herbivores in three populations sampled along the latitudinal gradient demonstrated that intensity of herbivory and its effects on plant fitness decreased towards the north. A reciprocal transplant experiment among the same three populations showed that herbivory affected the relative fitness of the three populations, but did not detect any evidence of local adaptation. Instead the southernmost population had the highest relative fitness at all three sites. A herbivore-removal experiment conducted in nine populations in an archipelago in northern Sweden demonstrated that insect herbivory strongly influenced among-population variation in reproductive output. However, variation in resistance was not related to differences in intensity of herbivory at this spatial scale. Taken together, the results demonstrate that resistance and tolerance to herbivory vary with latitude but in opposite directions, that intensity of herbivory is a major determinant of flowering and seed output, and that the strength of herbivore-mediated selection varies among populations in Lythrum salicaria. They further indicate that both physical disturbance regime and latitudinal variation in abiotic conditions may strongly influence the performance and abundance of perennial herbs because of their effects on interactions with specialized herbivores.

Disturbance at key areas used by wolves may have negative impacts on population dynamics and species recovery, becoming particularly critical in regions with high human activity, such as the Iberian Peninsula. This study evaluated the sources of human-related disturbances and ecological features in the Iberian wolf key areas, including feeding sites, highway crossing structures and, more exhaustively, breeding sites, using available data from camera trapping obtained in two areas within wolf range in Portugal. Differences at spatial and temporal level were observed in the use of these areas by wolves, other wildlife, domestic animals and humans. A decreasing use of these sites by wildlife with an increase in the use by domestic animals and humans was predicted. However, wild species shown to benefit from areas with intermediate human intervention. The expansion of traditional agriculture and fire mitigation measures seems to mitigate or revert the decreasing trends of wolves, mesocarnivores and wild ungulates; Locais de reprodução do lobo em territórios humanos: perceções sobre as características do habitat e fontes de perturbação para apoiar medidas de conservação RESUMO: Perturbações em áreas-chave ocupadas pelos lobos podem causar impactos negativos na dinâmica populacional e na recuperação desta espécie, tornando-se particularmente crítico em regiões com elevada presença humana, como na Península Ibérica. Este estudo avaliou as fontes de perturbações humanas e características ecológicas nas áreas-chave do lobo-ibérico, incluindo locais de alimentação, passagens de autoestradas e, mais exaustivamente, locais de reprodução, utilizando dados de armadilhagem fotográfica obtida em duas áreas de distribuição do lobo em Portugal. Diferenças espaciais e temporais foram observadas nas deteções de lobos, outros animais selvagens, animais domésticos e humanos. Foi prevista uma diminuição do uso desses locais pela fauna selvagem com um aumento do uso por animais domésticos e humanos. No entanto, espécies selvagens demonstraram beneficiar de áreas com intervenção humana intermédia. A expansão da agricultura tradicional e medidas de mitigação de incêndios mostraram desacelerar ou mesmo reverter as tendências decrescentes de lobos, mesocarnívoros e ungulados selvagens.

There are large gaps in our understanding how fish populations are affected by the anthropogenic noise and the alteration of habitat caused by the construction and operation of offshore wind farms. These issues are of great importance as the construction of offshore wind farms will increase all over the world in the near future. This thesis studies these effects with a focus on fish. The wind turbine foundations function as artificial reefs and are colonized by invertebrates, algae and fish. The epibenthic assemblages are influenced by factors such as hydrographical parameters, time of submergence, distance to natural hard bottom, material and texture (PAPER I, II). Once an epibenthic assemblage has been developed, fish utilize it for different ecosystem services such as food, shelter, and spawning and nursery area. Benthic and semi-pelagic species show a stronger response to the introduced foundation than pelagic species, as it is the bottom habitat that has mainly been altered (PAPER I, II). Pelagic species could be positively affected by the increased food availability - but it takes time and the effect is local. Construction noise like pile driving creates high levels of sound pressure and acoustic particle motion in the water and seabed. This noise induces behavioural reactions in cod (Gadus morhua) and sole (Solea solea). These reactions could occur up to tens of kilometres distance from the source (PAPER III). During power production, the wind turbines generate a broadband noise with a few dominating tones (PAPER IV, V), which are detectable by sound pressure sensitive fish at a distance of several kilometres even though intense shipping occurs in the area. Motion sensitive species will only detect the turbine noise at around a ten meter distance. Sound levels are only high enough to possibly cause a behavioural reaction within meters from a turbine (PAPER IV, V).
At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 3, 4 and 5: Manuscripts.