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1

MALETZ, JORG, and DAVID L. BRUTON. "The Beothuka terranova (Radiolaria) assemblage and its importance for the understanding of early Ordovician radiolarian evolution." Geological Magazine 142, no. 6 (November 2005): 711–21. http://dx.doi.org/10.1017/s0016756805001391.

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The radiolarian Beothuka terranova occurs in the Arenigian Didymograptellus bifidus Biozone (uppermost Lower Ordovician) of Spitsbergen (Svalbard), associated with a diverse and well-preserved radiolarian fauna. The presence of typical Cambrian spicular radiolarians associated with derived spherical forms shows a gradational faunal change from the Cambrian to the Ordovician. The genus and species Antygopora ordovicica n. gen. et n. sp. is described.
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Zhang, Yuandong, Bernd-D. Erdtmann, and Hongzhen Feng. "Tremadocian (Early Ordovician) graptolite biostratigraphy of China." Newsletters on Stratigraphy 40, no. 3 (December 22, 2004): 155–82. http://dx.doi.org/10.1127/0078-0421/2004/0040-0155.

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3

Retallack, Gregory J. "Ordovician Life on Land and Early Paleozoic Global Change." Paleontological Society Papers 6 (November 2000): 21–46. http://dx.doi.org/10.1017/s1089332600000693.

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Many Paleontologists share the opinion of McGhee (1996), who wrote “Prior to the Devonian, there was no terrestrial ecosystem to speak of. Some primitive plants precariously establishing a beachhead in protected coastal areas was about it. The interiors of the continents of the planet Earth were as barren as the rocky landscapes of Mars.” Thus, it was with trepidation that I reported paleosols containing trace fossils of early land animals in the late Ordovician, Juniata Formation, of Pennsylvania (Retallack and Feakes, 1987; Retallack, 1992a, 1992b, 1993). My late colleague, Jane Gray, engendered considerable debate by reporting Ordovician and Early Silurian spores like those of liverworts (Gray and Boucot, 1977; Gray, 1985). This spore, trace fossil and paleosol evidence for life on land in the Ordovician has remained controversial (Buatois et al., 1998; Shear, 1998), but evidence for Ordovician life on land has continued to accumulate. Especially important was discovery of myriapod trackways from mid-Ordovician (Llandeilian-Caradocian) Borrowdale Volcanics of the Lake District, England (Johnson et al., 1994). Abundant arthropod burrows and tracks, and a single body fossil of an euthycarcinoid in the fluvial-eolian Tumblagooda Sandstone of Western Australia (White 1990; McNamara and Trewin, 1993; Trewin and McNamara, 1995) are now thought to be late Ordovician in age (Iaksy et al., 1998). An enigmatic assemblage of arthropods and plants from a mid-Ordovician paleokarst in Tennessee (Caster and Brooks, 1956) is now thought to have been lacustrine (Gray, 1988a). The fossil record of Ordovician land plants also has improved with the discovery of possible megafossil mosses (Snigirevskaya et al. 1992), and possible late Ordovician trilete spores (Nøhr-Hansen and Koppelhus, 1998; Richardson 1988; Strother, 1991; Strother et al., 1996). But the most abundant evidence for Ordovician life on land remains fossil soils, now exploited by increasingly thorough and sophisticated studies (Retallack, 1985, 1992a, 1992b, 1993; Feakes et al., 1989; Driese and Foreman 1991, 1992a, 1992b; Driese et al., 1992, 1997; Mora et al., 1991, 1996; Mora and Driese, 1993; Yapp and Poths, 1992, 1994, 1996; Yapp, 1993, 1996). Mounting evidence from fossils and paleosols now presents an increasingly detailed view of Ordovician ecosystems on land.
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4

Sprinkle, James, and Thomas E. Guensburg. "Early radiation of echinoderms." Paleontological Society Papers 3 (October 1997): 205–24. http://dx.doi.org/10.1017/s1089332600000267.

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Echinoderms underwent a major two-part radiation that produced all of the major groups found in the fossil record between the Early Cambrian and the Middle Ordovician. A small initial radiation in the Early and Middle Cambrian produced about nine classes containing low-diversity members of the Cambrian Evolutionary Fauna. These were characterized by primitive morphology, simple ambulacral feeding structures, and the early development of a multiplated stalk or stem for attachment to skeletal fragments on a soft substrate. Several groups became extinct at the end of the Middle Cambrian, leaving the Late Cambrian as a gap of very low diversity in the fossil record of echinoderms with only four classes preserved and very few occurrences of complete specimens, mostly associated with early hardgrounds. The survivors from this interval re-expanded in the Early Ordovician and were joined by many newly evolved groups to produce a much larger radiation of more advanced, diverse, and successful echinoderms representing the Paleozoic Evolutionary Fauna on both hard and soft substrates. At least 17 classes were present by the Middle Ordovician, the all-time high point for echinoderm class diversity, and nearly all of the major ways-of-life (except for deep infaunal burrowing) had been developed. With the rise to dominance of crinoids, many less successful or archaic groups did not survive the Middle Ordovician, and echinoderm class diversity dropped further because of the mass extinction at the end of the Ordovician. This weeding-out process of other less-successful echinoderm groups continued throughout the rest of the Paleozoic, and only five classes of echinoderms have survived to the Recent from this early Paleozoic radiation.
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5

Runnegar, Bruce. "Origin and Early History of Mollusks." Notes for a Short Course: Studies in Geology 13 (1985): 17–32. http://dx.doi.org/10.1017/s027116480000107x.

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Fossil mollusks are common and conspicuous objects in shallow marine strata of Ordovician and younger ages, and so the broad features of the post-Cambrian history of the Mollusca have been known for many years. However, as all but one of the eight classes of the Mollusca had evolved by the beginning of the Ordovician, it is obvious that the pre-Ordovician history of the Mollusca holds the key to an understanding of the primary radiation of the phylum. It was with this view in mind, that John Pojeta, Jr. and I began our joint studies of Cambrian mollusks some 13 years ago.
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6

Norford, B. S., and M. P. Cecile. "Ordovician emplacement of the Mount Dingley Diatreme, Western Ranges of the Rocky Mountains, southeastern British Columbia." Canadian Journal of Earth Sciences 31, no. 10 (October 1, 1994): 1491–500. http://dx.doi.org/10.1139/e94-132.

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External and internal morphologies are well shown by a newly discovered diatreme that is exceptionally well exposed in a cirque within the north face of Mount Dingley. The diatreme contains abundant brecciated host rocks mixed with highly altered, fine-grained, light-green igneous fragments (minerals include muscovite, chlorite, quartz, carbonate, and some remnant K-feldspar). The diatreme cuts Lower Ordovician rocks of the McKay Group. Olistostromes and other volcaniclastic rocks that are directly associated with the diatreme are bevelled beneath a regional unconformity below the Upper Ordovician Beaverfoot Formation. Lower Ordovician gastropods are present just below the volcaniclastic rocks and within what appears to be a lens of sediment within one of the olistostrome beds. These occurrences indicate a mid-Early Ordovician time of intrusion, but there is the possibility that the pipe was emplaced later within the interval mid-Early to early Late Ordovician. In the Western Ranges, three other episodes of emplacement of diatremes have been documented previously as within the intervals early Middle to early Late Ordovician, latest Early Silurian to early Middle Devonian, and Late Permian. Many of the diatremes are broadly contemporaneous with widespread, but volumetrically small, Ordovician and Lower Paleozoic volcanic and intrusive rocks found throughout the Canadian Cordillera. These volcanic and intrusive rocks have been interpreted as evidence of continued Lower Paleozoic extensional tectonism and some are associated with large base-metal deposits.
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7

Blake, Daniel B., Forest J. Gahn, and Thomas E. Guensburg. "Two new early Asteroidea (Echinodermata) and early asteroid evolution." Journal of Paleontology 94, no. 4 (April 2, 2020): 734–47. http://dx.doi.org/10.1017/jpa.2020.7.

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AbstractAerliceaster nexosus n. gen. n. sp. (Echinodermata), one of the oldest of known asteroids, is based on six specimens from the Floian (Early Ordovician) Garden City Formation of Idaho, and Kolataster perplexus n. gen. n. sp. is based on two specimens from the Sandian (Late Ordovician) Mifflin Formation of Illinois. Although the asterozoan skeleton is subdivided into few ossicular categories, evolutionary derivations of all the categories are not fully established, and therefore published evaluations differ. Beginning with phylogenetic work placing asteroid ancestry within the Somasteroidea together with the new taxa described herein, aspects of early asteroid morphology are evaluated and ambiguities in need of further study identified. Uncertainties are considered to be founded in rapid early asterozoan diversification and the scanty fossil record.UUID: http://zoobank.org/b43d07cc-c8fb-4a84-bc6f-40aa6e0daea2
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8

Harland, W. Brian. "Chapter 14 Cambrian-Ordovician history." Geological Society, London, Memoirs 17, no. 1 (1997): 257–71. http://dx.doi.org/10.1144/gsl.mem.1997.017.01.14.

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Cambrian-Ordovician history is well documented in Svalbard with late Early Cambrian faunas and a range of Ordovician faunas to provide a basis for correlation. Not so extensive as Vendian, the rocks crop out in four areas: (i) only slightly deformed strata in the youngest Hecla Hoek (Oslobreen) Group in northeastern Svalbard yield especially rich Early to Mid-Ordovician faunas, (ii) The Hornsundian Geosyncline in south Spitsbergen with more variable facies and tectonic complications also exhibits Early Cambrian and Canadian strata, (iii) The Bjornoya succession reveals a marked hiatus between Vendian and Early and Mid-Ordovician strata, (iv) In western Svalbard the lack of Cambrian and Early Ordovician strata marks a distinct Mid Ordovician tectono-thermal event to be followed by ?Late Ordovician and Early Silurian strata. Indeed the above four areas correspond to distinct terranes which, having different affinities especially with areas in Greenland, give evidence of relatively distant areas and environments of formation. Evidence of Cambro-Ordovician volcanism is not recorded.Figure 14.1 lists the successions in the four areas mentioned according to the classification of rock units as abstracted from chapters 6, 7, 8, 9, 10 and 11, where their regional settings may be found. The outcrops are plotted on Fig. 14.2. The northeastern Svalbard strata are separated by Hinlopenstretet. This waterway divides Ny Friesland and Olav V Land in Spitsbergen from northwestern Nordaustlandet and occupies a syncline, but the successions although differently named are essentially continuous. In southern Spitsbergen the fjord Hornsund separates the successions to the south in Sorkapp Land
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9

QUINTON, PAGE C., LAURA SPEIR, JAMES MILLER, RAYMOND ETHINGTON, and KENNETH G. MACLEOD. "EXTREME HEAT IN THE EARLY ORDOVICIAN." PALAIOS 33, no. 8 (August 14, 2018): 353–60. http://dx.doi.org/10.2110/palo.2018.031.

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10

Vavrdová, Milada. "Early Ordovician provincialism in acritarch distribution." Review of Palaeobotany and Palynology 98, no. 1-2 (November 1997): 33–40. http://dx.doi.org/10.1016/s0034-6667(97)00023-7.

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11

CASAS, J. M. "Ordovician deformations in the Pyrenees: new insights into the significance of pre-Variscan (‘sardic’) tectonics." Geological Magazine 147, no. 5 (January 25, 2010): 674–89. http://dx.doi.org/10.1017/s0016756809990756.

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AbstractTwo deformational events which developed prior to the Variscan structures can be characterized in the Palaeozoic rocks of the Pyrenees: a Middle (?) Ordovician folding event and a Late Ordovician fracture episode. The Middle (?) Ordovician folding event gives rise to NW–SE- to N–S-oriented, metric- to hectometric-sized folds, without cleavage formation or related metamorphism. These folds can account for the deformation and uplift of the pre-Upper Ordovician (Cambro-Ordovician) sequence and for the formation of the Upper Ordovician unconformity. Ordovician folds control the orientation of the Variscan main-folding-phase minor structures, fold axes and intersection lineation in the Cambro-Ordovician sediments. The Late Ordovician fracture episode gave rise to normal faults affecting the lower part of the Upper Ordovician series, the basal unconformity and the underlying Cambro-Ordovician metasediments. Displacement of some of these faults diminishes progressively upwards of the series and tapers off in the upper part of the Upper Ordovician rocks, indicating that the faults became inactive during Late Ordovician times before deposition of the Ashgillian metasediments. Normal faults can be linked to the Upper Ordovician volcanic activity, which has been extensively described in the Pyrenees. The aforementioned deformation episodes took place after the Early Ordovician magmatic event, which gave rise to a large volume of plutonic rocks in the Pyrenees as in other segments of the European Variscides. This Middle Ordovician contractional event separated two extensional events in the Pyrenees from Early Ordovician to Silurian times. This event prevents us from assuming the existence of a continuous extensional regime through Ordovician and Silurian times, and suggests a more complex evolution of this segment of the northern Gondwana margin during the Ordovician.
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12

Goldberg, Samuel L., Theodore M. Present, Seth Finnegan, and Kristin D. Bergmann. "A high-resolution record of early Paleozoic climate." Proceedings of the National Academy of Sciences 118, no. 6 (February 1, 2021): e2013083118. http://dx.doi.org/10.1073/pnas.2013083118.

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The spatial coverage and temporal resolution of the Early Paleozoic paleoclimate record are limited, primarily due to the paucity of well-preserved skeletal material commonly used for oxygen-isotope paleothermometry. Bulk-rock δ18O datasets can provide broader coverage and higher resolution, but are prone to burial alteration. We assess the diagenetic character of two thick Cambro–Ordovician carbonate platforms with minimal to moderate burial by pairing clumped and bulk isotope analyses of micritic carbonates. Despite resetting of the clumped-isotope thermometer at both sites, our samples indicate relatively little change to their bulk δ18O due to low fluid exchange. Consequently, both sequences preserve temporal trends in δ18O. Motivated by this result, we compile a global suite of bulk rock δ18O data, stacking overlapping regional records to minimize diagenetic influences on overall trends. We find good agreement of bulk rock δ18O with brachiopod and conodont δ18O trends through time. Given evidence that the δ18O value of seawater has not evolved substantially through the Phanerozoic, we interpret this record as primarily reflecting changes in tropical, nearshore seawater temperatures and only moderately modified by diagenesis. Focusing on the samples with the most enriched, and thus likely least-altered, δ18O values, we reconstruct Late Cambrian warming, Early Ordovician extreme warmth, and cooling around the Early–Middle Ordovician boundary. Our record is consistent with models linking the Great Ordovician Biodiversification Event to cooling of previously very warm tropical oceans. In addition, our high-temporal-resolution record suggests previously unresolved transient warming and climate instability potentially associated with Late Ordovician tectonic events.
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13

Kröger, Björn, Franziska Franeck, and Christian M. Ø. Rasmussen. "The evolutionary dynamics of the early Palaeozoic marine biodiversity accumulation." Proceedings of the Royal Society B: Biological Sciences 286, no. 1909 (August 28, 2019): 20191634. http://dx.doi.org/10.1098/rspb.2019.1634.

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The early Palaeozoic Era records the initial biodiversification of the Phanerozoic. The increase in biodiversity involved drastic changes in taxon longevity, and in rates of origination and extinction. Here, we calculate these variables in unprecedented temporal resolution. We find that highly volatile origination and extinction rates are associated with short genus longevities during the Cambrian Period. During the Ordovician and Silurian periods, evolutionary rates were less volatile and genera persisted for increasingly longer intervals. The 90%-genus life expectancy doubled from 5 Myr in the late Cambrian to more than 10 Myr in the Ordovician–Silurian periods. Intervals with widespread ecosystem disruption are associated with short genus longevities during the Cambrian and with exceptionally high longevities during the Ordovician and Silurian periods. The post-Cambrian increase in persistence of genera, therefore, indicates an elevated ability of the changing early Palaeozoic marine ecosystems to sustainably maintain existing genera. This is evidence of a new level of ecosystem resilience which evolved during the Ordovician Period.
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14

VIZAN, HAROLDO, JOHN N. CARNEY, PETER TURNER, ROBERT A. IXER, MARK TOMASSO, ROBERT P. MULLEN, and PAUL CLARKE. "Late Neoproterozoic to Early Palaeozoic palaeogeography of Avalonia: some palaeomagnetic constraints from Nuneaton, central England." Geological Magazine 140, no. 6 (November 2003): 685–705. http://dx.doi.org/10.1017/s001675680300832x.

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Palaeomagnetic studies have been carried out on Neoproterozoic, Cambrian and Ordovician rocks in the Nuneaton inlier, England (52.5° N, 1.5° W). Three magnetic components were recognized, which provide a consistent structural and magnetic history of the inlier. Neoproterozoic volcaniclastic and intrusive rocks acquired a characteristic remanent magnetization (ChRM) dated at 603Ma. Late Ordovician rocks are represented by lamprophyre and diorite intrusions and their ChRMs were probably imprinted during their emplacement, at about 442 Ma. The Lower Cambrian sedimentary sequence of the Hartshill Sandstone Formation, which unconformably overlies the Neoproterozoic rocks and hosts the Ordovician intrusions, does not preserve a primary magnetization but shows the imprints of the Late Ordovician (442 Ma) remagnetization, as well as a probable end-Carboniferous remagnetization. Palaeolatitudes calculated for the late Neoproterozoic rocks and Ordovician intrusions are in good agreement with other palaeolatitudes calculated for Avalonia during those times. Both the late Neoproterozoic and Late Ordovician rocks additionally show ChRMs with declination anomalies indicating a large tectonic rotation of the Nuneaton area, possibly during one of the Caledonian phases of deformation affecting southern Britain.
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Brower, James C. "Flexible crinoids from the Upper Ordovician Maquoketa Formation of the northern midcontinent and the evolution of early flexible crinoids." Journal of Paleontology 75, no. 2 (March 2001): 370–82. http://dx.doi.org/10.1017/s0022336000018163.

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Three flexible crinoids occur in the Upper Ordovician Maquoketa Formation of Illinois, Iowa, and Minnesota:Protaxocrinus girvanensisRamsbottom, 1961,Clidochirus anebosnew species, andProanisocrinus oswegoensis(Miller and Gurley, 1894).Protaxocrinus girvanensisis also found in the Upper Ordovician of Scotland which indicates that the ocean was narrow enough to allow at least one crinoid species to cross the barrier. The Upper Ordovician of North America and Scotland also share many common crinoid genera. Both phenetic and cladistic methods result in similar phylogenies of flexible crinoids.Protaxocrinuswas derived from a cupulocrinid ancestor during the Middle Ordovician.Clidochirusevolved fromProtaxocrinusor its ancestral stock prior to the Richmondian of the Late Ordovician. The RichmondianProanisocrinusand later anisocrinids are most closely related toClidochirusor its immediate predecessor. Thus, three major lineages of flexible crinoids,Protaxocrinus(taxocrinid group),Clidochirus(icthyocrinid), andProanisocrinus(anisocrinids and homalocrinids), appeared during the Ordovician. Despite their rarity during the Ordovician, all three flexible lineages survived the Latest Ordovician extinction, whereas their more abundant and successful cupulocrinid ancestors were eliminated.
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Richardson, Jeffery G., and William I. Ausich. "Late Ordovician - Early Silurian cryptospore occurrences on Anticosti Island (Île d'Anticosti), Quebec, Canada." Canadian Journal of Earth Sciences 44, no. 1 (January 1, 2007): 1–7. http://dx.doi.org/10.1139/e06-100.

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Residues from Hirnantian (Upper Ordovician) and Aeronian (Lower Silurian) strata of Anticosti Island, Quebec, Canada, have yielded the oldest occurrence of terrestrial-based palynomorphs (cryptospores) from eastern North America. A low-abundance, low-diversity cryptospore assemblage containing specimens of Pseudodyadospora, Velatitetras, and alete monads has been recovered from the Velleda Member of the Ellis Bay Formation (Upper Ordovician, Hirnantian). The Gun River Formation (Lower Silurian, Aeronian) contains a low-abundance, low-diversity assemblage composed of specimens of Velatitetras, Laevolcancis, and alete monads. The occurrence of these assemblages provides evidence for early land plants in this part of North America during the Late Ordovician and Early Silurian, and documents the existence of a terrestrial-based flora during and after the Saharan (Late Ordovician) glaciation.
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Rasmussen, Christian M. Ø., Björn Kröger, Morten L. Nielsen, and Jorge Colmenar. "Cascading trend of Early Paleozoic marine radiations paused by Late Ordovician extinctions." Proceedings of the National Academy of Sciences 116, no. 15 (March 25, 2019): 7207–13. http://dx.doi.org/10.1073/pnas.1821123116.

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The greatest relative changes in marine biodiversity accumulation occurred during the Early Paleozoic. The precision of temporal constraints on these changes is crude, hampering our understanding of their timing, duration, and links to causal mechanisms. We match fossil occurrence data to their lithostratigraphical ranges in the Paleobiology Database and correlate this inferred taxon range to a constructed set of biostratigraphically defined high-resolution time slices. In addition, we apply capture–recapture modeling approaches to calculate a biodiversity curve that also considers taphonomy and sampling biases with four times better resolution of previous estimates. Our method reveals a stepwise biodiversity increase with distinct Cambrian and Ordovician radiation events that are clearly separated by a 50-million-year-long period of slow biodiversity accumulation. The Ordovician Radiation is confined to a 15-million-year phase after which the Late Ordovician extinctions lowered generic richness and further delayed a biodiversity rebound by at least 35 million years. Based on a first-differences approach on potential abiotic drivers controlling richness, we find an overall correlation with oxygen levels, with temperature also exhibiting a coordinated trend once equatorial sea surface temperatures fell to present-day levels during the Middle Ordovician Darriwilian Age. Contrary to the traditional view of the Late Ordovician extinctions, our study suggests a protracted crisis interval linked to intense volcanism during the middle Late Ordovician Katian Age. As richness levels did not return to prior levels during the Silurian—a time of continental amalgamation—we further argue that plate tectonics exerted an overarching control on biodiversity accumulation.
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18

Hall, Stuart A., and Ian Evans. "Paleomagnetic study of the Ordovician Table Head Group, Port au Port Peninsula, Newfoundland." Canadian Journal of Earth Sciences 25, no. 9 (September 1, 1988): 1407–19. http://dx.doi.org/10.1139/e88-135.

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A paleomagnetic study of the Ordovician Table Head Group in the Port au Port Peninsula of western Newfoundland reveals a simple two-component magnetization history comprising a reversely magnetized, stable southeasterly remanence with a, shallow to moderate inclination, and an unstable present-day overprint. Pole positions for the stable component, both with and without tectonic tilt correction, correspond with the Early to middle Ordovician pole positions for North America, suggesting this remanence is early. Although the nature of the geomagnetic field in the Ordovician is not well known, the polarity observed is consistent with that reported from other mid-Ordovician studies and appears to reflect a predominance of reverse polarity for this time interval. Evidence of significant rotation of any of the sites studied is absent, indicating that the continental margin in this region acted in an integral rather than a fragmented fashion during deformation.Previously published and new, but preliminary, results from the Early Ordovician St. George Group indicate the presence of two stable components of remanence. These components have similar south-southeasterly declinations but differ in inclination. The shallow to intermediate positive inclination component has a direction that is broadly compatible with Early Ordovician poles from North America. The shallow negative inclination component observed in rocks of similar age from other parts of western Newfoundland appears to be consistent with a later remagnetization of this unit.
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Smith, M. P., and J. S. Peel. "The age of the Danmarks Fjord Member, eastern North Greenland." Rapport Grønlands Geologiske Undersøgelse 132 (December 31, 1986): 7–13. http://dx.doi.org/10.34194/rapggu.v132.7957.

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Conodonts of late Early Ordovician age (late Canadian, early-middle Arenigian) are identified from the Danmarks Fjord Member of the Wandel Valley Formation at its type locality near the head of Danmark Fjord, eastern North Greenland. The identifications confirm recent suggestions of an Early Ordovician age for the member made on lithostratigraphic grounds, and refute earlier opinions that the dolomite was probably of Early Cambrian age.
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Cope, John C. W. "Early Ordovician Bivalve Radiations and Their Cause." Paleontological Society Special Publications 8 (1996): 83. http://dx.doi.org/10.1017/s247526220000085x.

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Cocks, L. Robin M., and Leonid E. Popov. "Early Ordovician brachiopods from south-west Wales." Proceedings of the Geologists' Association 130, no. 6 (December 2019): 677–90. http://dx.doi.org/10.1016/j.pgeola.2019.02.001.

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22

Hambrey, M. J. "The late Ordovician—Early Silurian glacial period." Palaeogeography, Palaeoclimatology, Palaeoecology 51, no. 1-4 (October 1985): 273–89. http://dx.doi.org/10.1016/0031-0182(85)90089-6.

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23

Pushkin, Victor I., and Leonid E. Popov. "Early Ordovician bryozoans from north-western Russia." Palaeontology 42, no. 1 (February 1999): 171–89. http://dx.doi.org/10.1111/1475-4983.00067.

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24

Sanford, Bruce V. "Stratigraphic and Structural Framework of Upper Middle Ordovician Rocks in the Head Lake-Burleigh Falls Area of South-Central Ontario." Géographie physique et Quaternaire 47, no. 3 (November 23, 2007): 253–68. http://dx.doi.org/10.7202/032956ar.

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ABSTRACT Field investigations in the Head Lake-Burleigh Falls area of south-central Ontario, that focused mainly on the Covey Hill(?), Shadow Lake, Gull River and Coboconk formations along the Paleozoic escarpment, provided clear evidence of faulting. Observed and inferred structural deformation, coupled with faciès changes within the Shadow Lake and lower Gull River, point to a succession of basement movements during the Phanerozoic. These range from Hadrynian-Early Cambrian, through Middle Ordovician to post-late Middle Ordovician times. Some of the earlier movements (Hadrynian-Early Cambrian to late Middle Ordovician) appear to be coincident with, and probably related to, plate tectonic events and the associated Taconian orogeny that were in progress along the southeastern margins of the North American continent. Post-Middle Ordovician block faulting, on the other hand, may have been triggered by any number of epeirogenic events related to late stage Taconian, Acadian or Alleghanian orogenies in Late Ordovician to Carboniferous times, or possibly to rifting associated with continental break-up and initiation of seafloor spreading processes in the early Mesozoic. Manuscrit révisé accepté le 5 août 1993
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Ingham, J. Keith, Gordon B. Curry, and Alwyn Williams. "Early Ordovician Dounans Limestone fauna, Highland Border Complex, Scotland." Transactions of the Royal Society of Edinburgh: Earth Sciences 76, no. 4 (1985): 481–513. http://dx.doi.org/10.1017/s0263593300010671.

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ABSTRACTA diverse, but sparsely distributed silicified fauna of over 30 taxa has been recovered from 7·5 tonnes of acid-etched Lower Ordovician Dounans Limestone from the Highland Border Complex, near Aberfoyle. The 13 trilobite taxa obtained include 3 new formally named species: Distazeris adoceta, Punka aetholiciocorus and Ischyrotoma stubblefieldi. Other elements, representing Carolinites, Illaenus, Goniotelina, Sycophantia, Kawina, Heliomeroides, Strotactinus, Ectenonotus, Ceratocephala and an indeterminate bathyurelline are described under open nomenclature although at least 4 are also probably new and 2 more may be conspecific with previously described species. The brachiopods include representatives of Archaeorthis, Nothorthis, Orthidium, ?Camerella, Idiostrophia, Orthambonites and 4 other indeterminate stocks. Gastropods tentatively assigned to Murchisonia, Subulites, Straparollina, Maclurites and Cyrtodiscus are described, as are bryozoans, an orthocone, crinoids, and Incertae sedis.This profoundly North American fauna is Late Canadian ( = mid Arenig) in age and equates with the Cassinian Stage: it is key evidence in showing that the Highland Border rocks are not part of the Dalradian Supergroup whose earliest deformations and metamorphism predate the Ordovician. The field relationships of the limestone, together with evidence from derived clasts in Middle Ordovician and Devonian sequences near Girvan in SW Scotland and at Stonehaven in eastern Scotland suggest that it forms part of a widespread sub-Old Red Sandstone carbonate sequence of Early Ordovician age beneath the northern Midland Valley.
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26

Li, Xing, and Mary Droser. "The development of Early Paleozoic shell concentrations: evidence from the Cambrian and Ordovician of the Great Basin." Paleontological Society Special Publications 6 (1992): 183. http://dx.doi.org/10.1017/s2475262200007437.

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Shell concentrations have constituted an important and conspicuous part of the stratigraphic record since the Early Cambrian. The paleontological and stratigraphic significance of shell beds is well understood, primarily from Mesozoic and Cenozoic examples. Lower Paleozoic fossil concentrations, however, have not received much attention. The Cambrian and Ordovician evolutionary radiations were two of the most significant events in the history of life and established the Cambrian and Paleozoic faunas respectively. In order to determine the effect of these radiations on the development of fossil accumulations, a systematic study of early Paleozoic shell beds was conducted in the Great Basin areas of California, Nevada, and Utah.In order to minimize taphonomic variations in original chemical and physical conditions, shell beds were compared from strata deposited in similar depositional environments from similar tectonic settings. Preliminary analysis of the shell beds from relatively pure carbonate facies and mixed carbonate and siliciclastic facies shows: 1) that shell concentrations became a significant stratigraphic feature in the later Early Cambrian; 2) the thickness and lateral extent of the shell beds increase from Early Cambrian to Middle Ordovician; 3) the abundance and internal complexity of the shell beds increase from Early Cambrian to Middle Ordovician; and 4) the Cambrian and Early Ordovician shell beds are primarily, if not exclusively, dominated by trilobites whereas the Middle Ordovician shell beds are dominated by brachiopods and ostracodes.These data show a temporal trend in the development of the early Paleozoic shell beds. The nature of the Cambrian and Ordovician shell beds differs qualitatively and quantitatively. There is an increase in physical scale, abundance, and internal complexity through time. The thickness and abundance of the trilobite beds increase through the Cambrian. Interestingly, although trilobites were still diverse and abundant, they did not commonly generate thick trilobite beds after the Late Cambrian. The early Middle Ordovician is a critical time in the development of early Paleozoic shell beds. A variety of monotaxic and polytaxic shell beds, including 6m thick composite beds, first appeared at this time. While the brachiopods and ostracodes generate laterally extensive, commonly monotaxic, shell beds, the gastropods and bryozoans only formed lenticular concentrations.
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27

Kröger, Björn, Matilde S. Beresi, and Ed Landing. "Early orthoceratoid cephalopods from the Argentine Precordillera (Lower-Middle Ordovician)." Journal of Paleontology 81, no. 6 (November 2007): 1266–83. http://dx.doi.org/10.1666/06-013.1.

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The Early and Middle Ordovician Orthocerida and Lituitida of Precordilleran Argentina are described, and their systematics and paleogeographic significance are revised. These cephalopods show a strong affinity to coeval faunas of North China, suggesting a location of the Precordillera at middle latitudes in the Southern Hemisphere east of the North China block and relatively close to the Gondwanan margin during the early Middle Ordovician. The descriptive terminology of characters of the septal necks, the position and shape of the siphuncule, and the shape of the connecting ring is improved. The distribution of these characters support an emendation of the Baltoceratidae, Sactorthoceratidae, and Proteoceratidae. Braulioceras n. gen. (Sactorthoceratidae) and Palorthoceras n. gen. (Orthoceratidae) are erected. The new species Braulioceras sanjuanense, Eosomichelinoceras baldisii, Gangshanoceras villicumense, and Rhynchorthoceras minor are proposed. Palorthoceras n. gen. from the Lower Ordovician Oepikodus evae Zone represents the earliest known orthocerid.
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28

Ruban, Dmitry. "Was there more space in the late Early Devonian for marine biodiversity to peak than in the early Late Ordovician?: A brief note." Annales g?ologiques de la Peninsule balkanique, no. 74 (2013): 1–8. http://dx.doi.org/10.2298/gabp1374001r.

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After the so-called ?Cambrian explosion?, marine biodiversity peaked either in the early Late Ordovician (as shown by the ?classical? curves based on the extensive palaeontological data compilation) or in the late Early Devonian (as shown by the ?innovative? curve based on the sampling standardization). The brief review of the modern plate tectonic, palaeoclimatic, and eustatic reconstructions demonstrates that shelves, which likely provided the main space for biotic radiation, shrank, concentrated in the tropics, and were better connected in the late Early Devonian than in the early Late Ordovician. The results of the present analysis permit to hypothesize that there was more (or the same) space for marine organisms to reach their maximum in their number in the early Late Ordovician relatively to the late Early Devonian. This is the only particular hypothesis, and the other extrinsic and intrinsic factors should be considered in further discussions.
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29

Fortey, R. A. "Early Ordovician trilobites from the Wandel Valley Formation, eastern North Greenland." Rapport Grønlands Geologiske Undersøgelse 132 (December 31, 1986): 15–25. http://dx.doi.org/10.34194/rapggu.v132.7959.

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A small trilobite fauna is described from the Wandel Valley Formation of Kronprins Christian land, eastern North Greenland. It has a specific composition identical to the fauna from the Catoche Formation, western Newfoundland, which is typical of the shallow water bathyurid biofacies of the eastern part of the Ordovician Laurentian palaeocontinent. The fauna is of early Ordovician age, trilobite Zone H, equivalent to the early Arenig.
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30

Chen, Ya-Dong, Shoufa Lin, and Cees R. van Staal. "Detrital zircon geochronology of a conglomerate in the northeastern Cape Breton Highlands: implications for the relationships between terranes in Cape Breton Island, the Canadian Appalachians." Canadian Journal of Earth Sciences 32, no. 2 (February 1, 1995): 216–23. http://dx.doi.org/10.1139/e95-018.

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Cape Breton Island has been interpreted as consisting of four zones of pre-Carboniferous rocks, but the relationships among them are controversial. To help resolve the controversy, we have dated detrital zircons from a conglomerate (part of the Cheticamp Lake Gneiss) in the Aspy terrane in the northeastern Cape Breton Highlands using the U–Pb method. The following ages were obtained: 462 ± 2 Ma (Middle Ordovician); ~492–488 Ma (6 ages; Early Ordovician); 552 ± 3 Ma (latest Precambrian–Early Cambrian); 620 ± 13 and 687 ± 4 Ma (Cadomian); and 809 ± 17, 1423 ± 10, 1462 ± 12, 1605 ± 14, 1644 ± 4, and 1911 ± 5 Ma (Proterozoic). The Middle Ordovician age sets a maximum age limit for deposition of the conglomerate, and supports an Ordovician–Silurian age for the Cheticamp Lake Gneiss. The Early Ordovician, latest Precambrian–Early Cambrian, and Cadomian ages match published ages from the Bras d'Or terrane (and its correlatives) and the Mira terrane (and its correlatives), and indicate provenance of the conglomerate from both terranes. They also indicate that the Bras d'Or and Mira terranes had been connected by the time of deposition of the conglomerate. The combination of the Cadomian and the Proterozoic ages is typical of parts of South America, supporting a suggestion that the Avalon Composite Terrane was derived from South America.
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31

Foote, Mike. "Survivorship analysis of Cambrian and Ordovician trilobites." Paleobiology 14, no. 3 (1988): 258–71. http://dx.doi.org/10.1017/s0094837300011994.

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Cohort analysis is used to investigate survivorship of trilobites originating during the Cambrian and Ordovician. Using a time-homogeneous branching model, it is estimated that trilobite genera and species originating during the Ordovician survived three times longer than Cambrian genera and species. Monte Carlo simulation of survivorship is used to show that (1) Cambrian and Ordovician survivorship are significantly different, (2) Ordovician cohorts conform more closely to the time-homogeneous model than do Cambrian cohorts, and (3) deviations from temporal homogeneity are more often produced by extraordinary extinction than by unusually slow turnover.When Early Ordovician cohorts are decomposed into genera within families that originated in the Cambrian versus the Ordovician, no evidence that Cambrian and Ordovician survivorship differences are clade-specific can be found. Ordovician genera of Cambrian affinity and of Ordovician affinity become extinct at similar rates.Some of the ultimate causes of these differences in survivorship include (1) taxonomic inconsistency, (2) greater environmental stability in the Ordovician, and (3) more highly structured ecosystems in the Ordovician that may have led to the weeding out of extinction-prone taxa.
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32

Peters, Shanan E., and William I. Ausich. "A sampling-adjusted macroevolutionary history for Ordovician-Early Silurian crinoids." Paleobiology 34, no. 1 (2008): 104–16. http://dx.doi.org/10.1666/07035.1.

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Temporal variation in sampling intensity and geologically controlled rates of fossil preservation distort macroevolutionary patterns in the fossil record. Here, we use a comprehensive, list-based compilation of taxonomically and stratigraphically vetted global crinoid genus occurrences to evaluate and correct for the effects of variable and incomplete sampling from the Ordovician through Early Silurian. After standardizing the number of occurrences or the number of biofacies used to estimate the stratigraphic ranges of genera and after adjusting rates of turnover to account for the incomplete preservation of true extinction and origination pulses, we find support for several important revisions to the macroevolutionary history of crinoids. First, in contrast to the uncorrected data, sample-standardized genus richness does not appear to increase by more than 20% after an abrupt Middle Ordovician (Harnagian) diversification. Second, the only significant short-term change in genus richness following the Harnagian increase is a ≥24% decline from the Rawtheyan to the Hirnantian. Third, volatility in rates of genus extinction is increased after adjusting for preservation and there remain significant peaks of extinction in the Rawtheyan, which marks the end-Ordovician extinction, and in the middle of the Early Silurian. Finally, significant increases in origination rates occur in the Early Silurian. These results reaffirm the importance of the end-Ordovician extinction for crinoids, but they also highlight the comparatively poorly sampled Early Silurian as a time of turnover among crinoids.Crinoid genus extinction rates are positively correlated with area-weighted rates of sedimentary package truncation, suggesting that extinction may have been controlled by physical environmental changes, such as the contraction of unique epicontinental sea habitats. The lack of a correlation between genus origination and sedimentary package initiation reinforces this hypothesis and suggests that other factors, such as evolutionary innovations and biotic interactions during the Ordovician radiation, may have been more important in controlling the diversification of crinoids.
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33

Schmitz, Ulrich. "Sequence stratigraphy of the NE Spanish Middle Cambrian to Early Ordovician section." Zeitschrift der Deutschen Gesellschaft für Geowissenschaften 157, no. 4 (December 1, 2006): 629–46. http://dx.doi.org/10.1127/1860-1804/2006/0157-0629.

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34

Bock, B., S. M. McLennan, and G. N. Hanson. "The Taconian orogeny in southern New England: Nd-isotope evidence against addition of juvenile components." Canadian Journal of Earth Sciences 33, no. 12 (December 1, 1996): 1612–27. http://dx.doi.org/10.1139/e96-122.

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Nd-isotope data for pre-Taconian (meta)sedimentary and igneous rocks, syn-Taconian (meta)sedimentary rocks, and Late Ordovician–Silurian plutonic rocks indicate that the Ordovician Taconian orogeny did not add significant amounts of juvenile crust to the Laurentian margin in southern New England. Nd-isotope compositions of Grenvillian crust and Late Proterozoic to Early Cambrian rift sediments range from εNd of −3.1 to −6.6 at 450 Ma. Sedimentary rocks deposited during the Cambrian and the early Middle Ordovician, which represent the drift stage of Laurentia, and earliest Taconian sedimentary rocks show more negative εNd(450 Ma), with a range from −11.7 to −13.3. Sedimentary rocks deposited in response to the Taconian orogeny have uniform εNd(450 Ma) values of about −8. Middle to Late Ordovician and Permian plutonic rocks from southwestern Connecticut have εNd(450 Ma) values of −2 to −5, which indicates that these rocks contain older crustal components. Rocks with juvenile Nd characteristics are the early Paleozoic Maltby Lake Volcanics (εNd(450 Ma) +8) from southwestern Connecticut, and Middle Ordovician igneous samples from the Hawley Formation (εNd(450 Ma) +6 to −0.6) in Massachusetts.
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35

Oterdoom, W. Heiko, Mike A. Worthing, and Mark Partington. "Petrological and Tectonostratigraphic Evidence for a Mid Ordovician Rift Pulse on the Arabian Peninsula." GeoArabia 4, no. 4 (October 1, 1999): 467–500. http://dx.doi.org/10.2113/geoarabia0404467.

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ABSTRACT During late Early Ordovician times an increase in the rate of subsidence in the Ghaba Salt Basin and western South Oman Salt Basin is suggested by the thick sequence of continental clastics of the Ghudun Formation. After a phase of rift-shoulder uplift and erosion, related to a renewed pulse of extension which may have initiated diapiric growth of salt structures in the Ghaba Salt Basin, sedimentation resumed again in the Mid Ordovician. During this period, the center of deposition shifted to the Saih Hatat area in North Oman. This paper documents seismic and well data, field investigations and petrological study of potassic mafic rocks from the Huqf area which were intruded in the eastern side of the Ghaba Salt Basin. A Mid Ordovician age of 461 ± 2.4 million years has been established for these rocks by the Argon-Argon step heating method. Analogy with the petrology and setting of similar potassic mafic rocks from the Rio Grande Rift in the western United States of America suggests that they were intruded into the shoulder of an intra-continental rift. The data provide the first clear evidence of a pulse of rift-shoulder uplift in the Huqf area during the Mid Ordovician. The 3-kilometer-thick Mid to Late Ordovician clastic sediments of the Amdeh Formation in North Oman, together with the occurrence of abnormally thick sedimentary sequences and volcanics in the Tabas Graben in Iran, are consistent with a period of break-up of eastern Gondwana. Together, the Ghaba-Saih Hatat and Tabas Basins are considered to be part of a failed rift arm. These observations further improve our regional knowledge of the Early to Late Ordovician tectonic setting of Oman and will assist in unlocking the hydrocarbon potential of classical rift-related structures consisting of early-rift Early Ordovician sand-prone reservoirs sealed by syn-rift Mid to Late Ordovician marine shales.
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36

Hersi, O. Salad, G. S. Nowlan, and D. Lavoie. "A revision of the stratigraphic nomenclature of the Cambrian–Ordovician strata of the Philipsburg tectonic slice, southern Quebec." Canadian Journal of Earth Sciences 44, no. 12 (December 1, 2007): 1775–90. http://dx.doi.org/10.1139/e07-041.

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The Philipsburg tectonic slice is bounded to the west by a northeast–southwest-trending thrust fault (Logan’s Line) and preserves 10 formations of Middle (?) to Late Cambrian (Milton, Rock River, and Strites Pond formations), Early Ordovician (Wallace Creek, Morgan Corner, Hastings Creek, and Naylor Ledge formations), and early Middle Ordovician (Luke Hill, Solomons Corner, and Corey formations) age. The strata were previously assigned to the Philipsburg Group. Early correlations between the Philipsburg succession and coeval strata of the St. Lawrence Platform were mainly based on sparse macrofauna and inferred stratigraphic position. Unconformities at the Cambrian–Ordovician and Early Ordovician – Middle Ordovician boundaries occurring in autochthonous St. Lawrence Platform and the allochthonous Philipsburg succession (Philipsburg tectonic slice) highlight new stratigraphic interpretations between the inner-shelf (St. Lawrence Platform) and the outer-shelf (Philipsburg) successions. The succession in the Philipsburg tectonic slice is divided into three new groups. The Middle (?) to Upper Cambrian Missisquoi Group (new) includes the Milton, Rock River, and Strites Pond formations. The upper boundary of the Missisquoi Group is defined by the upper unconformable contact between the Upper Cambrian Strites Pond Formation and overlying Lower Ordovician Wallace Creek Formation. The Missisquoi Group correlates with the Potsdam Group of the St. Lawrence Platform. The Lower Ordovician School House Hill Group (new) includes the Wallace Creek, Morgan Corner, Hastings Creek, and Naylor Ledge formations. The upper boundary of this group is marked by a regionally extensive unconformity at the top of the Naylor Ledge Formation and correlates with the younger Beekmantown-topping unconformity. The School House Hill Group is correlative with the lower to upper part of the Beekmantown Group (Theresa Formation and the Ogdensburg Member of the Beauharnois Formation) of the St. Lawrence Platform. The Middle Ordovician Fox Hill Group (new) consists of the Luke Hill, Solomons Corner, and Corey formations. This group correlates with the uppermost part of the Beekmantown Group (Huntingdon Member of the Beauharnois Formation and the Carillon Formation).
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37

Ausich, William I., and Bradley Deline. "Macroevolutionary transition in crinoids following the Late Ordovician extinction event (Ordovician to Early Silurian)." Palaeogeography, Palaeoclimatology, Palaeoecology 361-362 (November 2012): 38–48. http://dx.doi.org/10.1016/j.palaeo.2012.07.022.

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38

Tollerton, Victor P. "Summary of a revision of New York State Ordovician eurypterids: implications for eurypterid palaeoecology, diversity and evolution." Transactions of the Royal Society of Edinburgh: Earth Sciences 94, no. 3 (September 2003): 235–42. http://dx.doi.org/10.1017/s026359330000064x.

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ABSTRACTThe record of Ordovician Eurypterida from New York State, USA, is shown to be largely false. Twenty-nine species in 17 genera are here recognised as pseudofossils, reducing by more than 75% the total number of named Ordovician eurypterid taxa. Consequently, 10 families now have their first occurrence either later in the Ordovician or in the Early Silurian. The implications for eurypterid palaeoecology, diversity and evolution are not as straightforward as would be expected from such a drastic taxonomic revision. All Ordovician eurypterids are now known to occur in shallow-water, near-shore shales or fine-grained carbonates. Diversity measures indicate that the end-Ordovician extinction event appears to have had less effect on eurypterids than previously known, and their turnover is level in the Ordovician.
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39

Fortey, R. A., R. M. Owens, and A. W. A. Rushton. "The palaeogeographic position of the Lake District in the early Ordovician." Geological Magazine 126, no. 1 (January 1989): 9–17. http://dx.doi.org/10.1017/s0016756800006105.

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AbstractThe early Ordovician was a time of maximum continental separation and hence a time when faunal evidence can be used to assess palaeogeography in a critical way. We summarize the known trilobite occurrences (18 genera) from the Arenig–Llanvirn of the Lake District, and record some genera for the first time. Maps of the distribution of some of these forms are given. All genera except Cyclopyge were confined to the Gondwana continent at the time, and some are known from many localities; and two species are widespread in England, Wales, France, Iberia and Bohemia. The fauna is entirely distinct from those of Scandinavia and North America. All the palaeontological evidence points to the Lake District being adjacent to Ordovician Gondwana. In the earlier Ordovician it is not reasonable to suggest that the Iapetus Ocean lay to the south of the Lake District as did Allen (1987).
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40

Nielsen, Arne Thorshøj. "A review of Ordovician agnostid genera (Trilobita)." Transactions of the Royal Society of Edinburgh: Earth Sciences 87, no. 4 (1996): 463–501. http://dx.doi.org/10.1017/s0263593300018150.

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ABSTRACTThe definition and scope of Ordovician genera of agnostid trilobites are discussed, and the generic affiliation of Ordovician species and well-illustrated material referred to in open nomenclature is assessed. Some 198 Ordovician species plus 25 species of uncertain latest Cambrian/earliest Ordovician age are enumerated, representing 24 genera and subgenera, of whichGeragnostus (Novoagnostus)n.subg. andLotagnostus (Semagnostus)n.subg. are new.Geragnostellais ranked as a subgenus ofGeragnostus. The species ofArthrorhachisare tentatively divided among atardusspecies group and anelspethispecies group. Species ofDividuagnostusare divided among asensu strictogroup and an early species group.PseudorhaptagnostusLermontova, 1951 is restored as senior synonym ofNeoagnostusKobayashi, 1955. It is concluded that Ordovician agnostids were largely adapted to temperate and colder water habitats.
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41

Blake, Daniel B. "Two Late Ordovician asteroids (Echinodermata) with characters suggestive of early ophiuroids." Journal of Paleontology 81, no. 6 (November 2007): 1476–85. http://dx.doi.org/10.1666/05-130.1.

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Jugiaster n. gen. is based on the Upper Ordovician asteroid (Echinodermata) Petraster speciosus; in addition, a second Upper Ordovician occurrence, Phyrtosaster casteri, new gen. and sp., is described. Both species exhibit mouth frame characters suggestive of those long considered typical of early ophiuroids, thereby demonstrating the adaptive versatility and complexity of asterozoan diversification. Jugiaster specimens also exhibit delicate ambulacral column articulation. The new fossils indicate the need for much further study before early asterozoan history will be well understood.
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42

ERNST, ANDREJ, OLGA K. BOGOLEPOVA, BERNHARD HUBMANN, ELENA YU GOLUBKOVA, and ALEXANDER P. GUBANOV. "Dianulites (Trepostomata, Bryozoa) from the Early Ordovician of Severnaya Zemlya, Arctic Russia." Geological Magazine 151, no. 2 (June 5, 2013): 328–38. http://dx.doi.org/10.1017/s0016756813000150.

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AbstractTrepostome bryozoan Dianulites borealis Astrova, 1965, the earliest known member of this genus, has been identified from the Early Ordovician of Severnaya Zemlya, Arctic Russia. This species developed hemispherical colonies which indicate that it lived on a relatively soft substrate with moderately low rates of sedimentation and erosion. The new record from Severnaya Zemlya expands the palaeogeographical distribution of Dianulites, known before from the Early Ordovician of Novaya Zemlya, Arctic Russia.
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43

Whalen, Joseph B. "The Topsails igneous suite, western Newfoundland: an Early Silurian subduction-related magmatic suite?" Canadian Journal of Earth Sciences 26, no. 12 (December 1, 1989): 2421–34. http://dx.doi.org/10.1139/e89-207.

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The Topsails igneous suite contains several Late Ordovician to Early Silurian volcanic and intrusive sequences, which overlie and intrude Early to Middle Ordovician oceanic and arc rocks. The oldest components of this suite may represent calc-alkaline, continental-arc magmatism. The younger components are bimodal, with felsic compositions vastly predominating, and include a major (> 2200 km2) alkaline (A-type) granite complex. These felsic components have similarities to peralkaline suites formed in unusual subduction-related settings. Younger mafic components resemble within-plate basalts emplaced in a continental setting.Silurian magmatic activity in the Canadian Appalachians is widespread, includes diverse magmatic types, and has contrasting metamorphic and tectonic overprinting, even in contiguous areas. These features and the probability of major post-Silurian displacements in the orogen render correlation and interpretation difficult. Tectonic models that consider basin closure and major plate movements to be complete by Middle Ordovician time fail to adequately explain the Silurian activity. Available data best fit a model that relates Late Ordovician to Silurian magmatic activity to the opening and closing of small, discontinuous basins, portions of which may have been floored by oceanic crust.
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44

RUSHTON, ADRIAN, MARK WILLIAMS, NGUYEN DUC PHONG, TOSHIFUMI KOMATSU, DAVID SIVETER, JAN ZALASIEWICZ, DINH CONG TIEN, NGUYEN VIET HIEN, NGUYEN HUU MANH, and GENGO TANAKA. "Early Ordovician (Tremadocian and Floian) graptolites from the Than Sa Formation, northeast Vietnam." Geological Magazine 155, no. 7 (April 24, 2017): 1442–48. http://dx.doi.org/10.1017/s0016756817000310.

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AbstractThe lower Palaeozoic marine succession of NE Vietnam accumulated on the South China plate. Despite historical works dating to French colonial times, the stratigraphy and palaeontology of the succession is poorly constrained. Chief amongst the lower Palaeozoic lithostratigraphical divisions is the Than Sa Formation, a c. 1200 m thick succession of clastic rocks of Cambrian and early Ordovician age. Newly collected graptolites (including Rhabdinopora? sp. and Tetragraptus approximatus) from the upper part of the formation identify strata assignable to the Tremadocian and Floian stages of the Lower Ordovician. The same succession also probably records the Cambrian–Ordovician boundary. Our analysis identifies one or more intervals of graptolite-bearing laminated mudstones in the upper part of the Than Sa Formation that may be widespread in NE Vietnam during the early Tremadocian.
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45

Harper, David A. T., Borja Cascales-Miñana, and Thomas Servais. "Early Palaeozoic diversifications and extinctions in the marine biosphere: a continuum of change." Geological Magazine 157, no. 1 (December 3, 2019): 5–21. http://dx.doi.org/10.1017/s0016756819001298.

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AbstractA review of biodiversity curves of marine organisms indicates that, despite fluctuations in amplitude (some large), a large-scale, long-term radiation of life took place during the early Palaeozoic Era; it was aggregated by a succession of more discrete and regionalized radiations across geographies and within phylogenies. This major biodiversification within the marine biosphere started during late Precambrian time and was only finally interrupted in the Devonian Period. It includes both the Cambrian Explosion and the Great Ordovician Biodiversification Event. The establishment of modern marine ecosystems took place during a continuous chronology of the successive establishment of organisms and their ecological communities, developed during the ‘Cambrian substrate revolution’, the ‘Ordovician plankton revolution’, the ‘Ordovician substrate revolution’, the ‘Ordovician bioerosion revolution’ and the ‘Devonian nekton revolution’. At smaller scales, different regional but important radiations can be recognized geographically and some of them have been identified and named (e.g. those associated with the ‘Richmondian Invasion’ during Late Ordovician time in Laurentia and the contemporaneous ‘Boda event’ in parts of Europe and North Africa), in particular from areas that were in or moved towards lower latitudes, allowing high levels of speciation on epicontintental seas during these intervals. The datasets remain incomplete for many other geographical areas, but also for particular time intervals (e.g. during the late Cambrian ‘Furongian Gap’). The early Palaeozoic biodiversification therefore appears to be a long-term process, modulated by bursts in significant diversity and intervals of inadequate data, where its progressive character will become increasingly clearer with the availability of more complete datasets, with better global coverage and more advanced analytical techniques.
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Guensburg, Thomas E., and James Sprinkle. "Environmental controls of rapidly diversifying echinoderms during the Early Paleozoic." Paleontological Society Special Publications 6 (1992): 114. http://dx.doi.org/10.1017/s2475262200006742.

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Echinoderms were among the earliest phyla to exploit global environmental shifts leading to the rise of the Paleozoic Evolutionary Fauna during the Ordovician. Echinoderms including crinoids and rhombiferans diversified more rapidly and actually precede other groups such as bryozoans, corals, and many articulate brachiopods in the record. We argue that preadaptive morphologic strategies among the echinoderms coupled with newly available and widespread living space led to this success.Sessile attached echinoderms, especially crinoids and some blastozoans, dominate echinoderm faunas generally throughout the Paleozoic. These organisms were rheophilic suspension feeders with elongate stalked morphologies capable of attaining high feeding levels above competitors. However, this morphology required firm attachment. Suitable sites were relatively rare in Early and especially Middle Cambrian time when shelf and slope sedimentation was dominated by fine siliciclastics and nonbiogenic carbonates resulting in extensive soft substrates. Sea level was relatively low during the Middle Cambrian, restricting shallow shelf living space. Stalked echinoderms then lived attached to scattered bioclastic fragments but had low diversity and were a relatively minor component of the faunal record. Mode of occurrence and morphologic features suggest they were generally unable to cope with soft and unstable substrates. Nearshore shelf sedimentation styles changed during the Late Cambrian and Early Ordovician to bioclastic-dominated carbonates and organic buildups. These were often subject to intermittent storm events and rapid submarine cementation of the seafloor because of changes in global atmospheric and seawater chemistry. This resulted in formation of hardgrounds on intraformational conglomerates, grainstones, and buildups. Gradual sea level rise beginning in the Late Cambrian and continuing through the Early Ordovician was a major contributing cause for these changes. The widespread lithified substrates provided ideal attachment sites for the preadapted echinoderms and greatly expanded the total habitable area, resulting in rapidly increasing diversity.Among crinoids, disparid inadunates were the first to extensively exploit this situation and they dominated throughout the Early Ordovician. Attached blastozoans were also common both in the Late Cambrian and Early Ordovician. Later, many camerate and cladid inadunate crinoids and a few blastozoans developed attachment styles suitable to soft substrates, continuing and expanding this diversification. Some previous authors have misidentified life modes of these Cambro-Ordovician echinoderms because they used forced senarios based on phylogenetic analysis alone. In contrast, we argue that life modes and paleoecologic settings are basic field-derived data that should be used along with morphologic character analysis to determine phylogenetic relationships. Both the fossils and their environmental settings must be studied together to reconstruct the adaptive history and phylogeny of a rapidly-radiating group.
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47

Deline, Bradley, and William I. Ausich. "Testing the plateau: a reexamination of disparity and morphologic constraints in early Paleozoic crinoids." Paleobiology 37, no. 2 (2011): 214–36. http://dx.doi.org/10.1666/09063.1.

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Studies of crinoid morphology have been pivotal in understanding the constraints on the range of morphology within a clade as well as the patterns of disparity throughout the Phanerozoic. Newly discovered and described faunas and recent study of early Paleozoic crinoid diversity provide an ideal opportunity to reanalyze Ordovician through Early Silurian crinoid disparity with more complete taxonomic coverage and finer stratigraphic resolution. Using the coarse stratigraphic binning of Foote (1999), the updated morphologic data set has a similar disparity pattern to those previously reported for the early Paleozoic. However, with the more resolved stratigraphic binning used by Peters and Ausich (2008), a significant difference exists between the original and current data sets. Both data sets have a pronounced disparity high during the late Middle Ordovician. However, the updated disparity curve has a much higher initial disparity during the Early Ordovician and a pronounced rise in disparity during the Silurian recovery. Examination of differential sampling, proportions of the crinoid orders through time, and methods of coding characters indicate these factors have little effect on the pattern of crinoid disparity. The Silurian morphospace expansion occurs primarily within disparids and coincides with the origination of the myelodactylids. These findings corroborate the rapid expansion of morphospace during the Ordovician. However, crinoid disparity did not remain static and, although less frequent than during the initial radiation, new body plans evolved following the Ordovician Extinction (e.g., the myelodactylids). These results are consistent with the hypothesis of ecology constraining the limits on morphologic disparity at the class level.
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48

KRÖGER, BJÖRN, and AMELIA PENNY. "EARLY–MIDDLE ORDOVICIAN SEASCAPE-SCALE AGGREGATION PATTERN OF SPONGE-RICH REEFS ACROSS THE LAURENTIA PALEOCONTINENT." PALAIOS 35, no. 12 (December 17, 2020): 524–42. http://dx.doi.org/10.2110/palo.2020.039.

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ABSTRACT During the late Cambrian–Early Ordovician interval the predominant non-microbial reef builders were sponges or sponge-like metazoans. The lithological and faunal composition of Cambro-Ordovician sponge-dominated reefs have previously been analyzed and reviewed. Here we take the relationship between reef aggregation pattern at reef to seascape scale into account, and look for changes during the Early–Middle Ordovician interval, in which metazoans became dominant reef builders. In a comparison of sponge-rich reefs from eight sites of the Laurentia paleocontinent three different seascape level reef growth patterns can be distinguished: (1) mosaic mode of reef growth, where reefs form a complex spatial mosaic dependent on hard substrate; (2) episodic mode, where patch reefs grew exclusively in distinct unconformity bounded horizons within non-reefal lithological units that have a much larger thickness; and (3) belt-and-bank mode, where reefs and reef complexes grew vertically and laterally as dispersed patches largely independent from truncation surfaces. The distinct modes of growth likely represent specific reef forming paleocommunities, because they differ in content and abundance of skeletal metazoan framebuilders, bioturbation intensity of non-skeletal reef sediment matrix, and in association of reef growth with underlying hard substrate. We suggest, based on a review of Laurentian reef occurrences, that the mosaic mode dominated in Early Ordovician strata and that the dominance shifted toward the belt and bank mode from Middle Ordovician strata onward.
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Carrera, Marcelo G., and J. Keith Rigby. "Biogeography of Ordovician sponges." Journal of Paleontology 73, no. 1 (January 1999): 26–37. http://dx.doi.org/10.1017/s0022336000027517.

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Sponges have an unrealized potential importance in biogeographic analysis. Biogeographic patterns determined from our analysis of all published data on distribution of Ordovician genera indicate Early Ordovician sponge faunas have relatively low diversity and are completely dominated by demosponges. Early Ordovician (Ibexian) faunas are characterized by the widespread co-occurrence ofArchaeoscyphiaand the problematicCalathium.This association is commonly found in biohermal structures. Middle Ordovician faunas show an increase in diversity, and two broad associations are differentiated: Appalachian faunas (including Southern China and the Argentine Precordillera) and Great Basin faunas.Late Ordovician faunas show important changes in diversity and provincialism. Hexactinellid and calcareous sponges became important and new demosponge families appeared. Four Mohawkian-Cincinnatian associations are recognized here, including: 1) Midcontinent faunas; 2) Baltic faunas; 3) New South Wales faunas; and 4) Western North American (California and Alaska) faunas. However, two separate biogeographic associations are differentiated based on faunal differences. These are a Pacific association (western North American and New South Wales) and an Atlantic association (Midcontinent Laurentia and Baltica).Distribution of sponge genera and migration patterns are utilized to consider paleogeographic dispositions of the different continental plates, climatic features, and oceanic currents. Such an analysis points to close paleogeographic affinities between the Argentine Precordillera and Laurentian Appalachian faunas. However, significant endemicity and the occurrence of extra-Laurentian genera suggest a relative isolation of the Precordillera terrane during the Late Ibexian-Whiterockian. The study also shows a faunal migration from the Appalachian region to South China during the Middle Ordovician and the migration of faunas from Baltica to Laurentia in the Late Ordovician. The occurrence of Laurentian migrants in New South Wales during the Late Ordovician could be related to inferred oceanic current circulation between these two areas, although other paleogeographic features may be involved.
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Laurie, John R., and Clive Burrett. "Biogeographic significance of Ordovician brachiopods from Thailand and Malaysia." Journal of Paleontology 66, no. 1 (January 1992): 16–23. http://dx.doi.org/10.1017/s0022336000033448.

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Early Ordovician brachiopods Spanodonta floweri (Cooper) and Aporthophyla tianjingshanensis? Fu from peninsular Thailand and the Langkawi Islands, Malaysia, and described and offer important confirmatory evidence for the Ordovician juxtaposition of the Shan-Thai terrane, the North China terrane, and Western Australia.
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