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1

Devaki, K., K. Senthilkumar, Pathan Nasrullahkhan, and M. G. Jayathangaraj. "Ancylostomiasis in Dhole Cuon alpinus." Zoos' Print Journal 21, no. 1 (December 21, 2005): 2141–42. http://dx.doi.org/10.11609/jott.zpj.1404.2141-2.

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Scala, Christopher, Katia Ortiz, Alexandra Nicolier, and Alexandra Briend-Marchal. "LYMPHOCYTIC LEUKEMIA IN A CAPTIVE DHOLE (CUON ALPINUS)." Journal of Zoo and Wildlife Medicine 44, no. 1 (March 2013): 204–7. http://dx.doi.org/10.1638/1042-7260-44.1.204.

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ALLWIN, BOON, R. THIRUMURUGAN, M. PALANIVEL RAJAN, and N. R. SENTHIL. "Trypanosomosis in a captive wild dog (Cuon alpinus)." VETERINARY SCIENCE RESEARCH JOURNAL 6, no. 1 (April 15, 2015): 67–70. http://dx.doi.org/10.15740/has/vsrj/6.1/67-70.

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Mothilal, B. M., and M. Eric Ramanujam. "First photographic record of the Dhole Cuon alpinus (Mammalia: Carnivora: Canidae) from the Sirumalai Hills in Tamil Nadu, India." Journal of Threatened Taxa 12, no. 10 (July 26, 2020): 16373–76. http://dx.doi.org/10.11609/jott.5959.12.10.16373-16376.

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The Asiatic Wild Dog has a wide global range covering Central, South and South-East Asia. It has been recorded over most of the Indian subcontinent except for the deserts of western India and Eastern Ghats of Tamil Nadu. We hereby provide photographic evidence of the Asiatic Wild Dog Cuon alpinus from Khandige Estate in Sirumalai.
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Cervera, José. "Nuevos hallazgos de carnivoros en el complejo de yacimientos mesopleistocenos de La Sierra de Atapuerca, Burgos." Spanish Journal of Palaeontology 7, no. 3 (August 10, 2022): 21. http://dx.doi.org/10.7203/sjp.25011.

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Se informa de nuevos hallazgos de restos de Carnivoros en el complejo de yacimientos de la Sierra de Atapuer­ca. Se aiiaden dos nuevos taxa: Cuon alpinus europaeus para Trinchera-Galeria y Lynx pardina spelaea para la Sima de los Huesos. Por otra parte se describen nuevos restos de Canis lupus spp. de Trinchera-Galeria y Panthera leo de ambos yacimientos.
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Bashir, Tawqir, Tapajit Bhattacharya, Kamal Poudyal, Manjari Roy, and Sambandam Sathyakumar. "Precarious status of the Endangered dhole Cuon alpinus in the high elevation Eastern Himalayan habitats of Khangchendzonga Biosphere Reserve, Sikkim, India." Oryx 48, no. 1 (October 15, 2013): 125–32. http://dx.doi.org/10.1017/s003060531200049x.

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AbstractDuring 2008–2010 we investigated the ecology of the Endangered dhole or wild dog Cuon alpinus in Khangchendzonga Biosphere Reserve in the Eastern Himalaya in Sikkim, India. We conducted camera trapping (n = 71 sites, 6,278 camera-days) and sign surveying along trails (n = 24; 629.43 km of effort) to assess the relative abundance, distribution and activity pattern of the dhole. Morphological characteristics evident in the 61 camera-trap photographs indicate that the dhole population in the Reserve may be the rare and genetically distinct subspecies C. alpinus primaevus. We detected dholes over a wide elevation range (2,501–4,100 m) that encompassed the upper temperate, subalpine, and alpine scrub zones. Dholes were diurnal, with peak activity at 08.00–10.00. Analysis of 41 scats indicated a diet comprising mainly mountain ungulates, rodents and pikas Ochotona sp. Although the frequency of occurrence of rodents was highest (32%) in the scats, 98.7% of the total biomass consumed was of mountain ungulates. Historical reports (1888–1894) indicated that the dhole was formerly abundant in Sikkim but was hunted to meet the high demand for its alleged medicinal properties. With no information on the status of the dhole in Sikkim for over a century, our study suggests that the species is now rare in the Reserve. To aid the conservation of the dhole and its main ungulate prey species extensive research and monitoring are required in the Reserve and elsewhere in the Eastern Himalaya.
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7

Thinley, Phuntsho, Jan F. Kamler, Sonam W. Wang, Kinzang Lham, Ute Stenkewitz, and David W. Macdonald. "Seasonal diet of dholes (Cuon alpinus) in northwestern Bhutan." Mammalian Biology 76, no. 4 (July 2011): 518–20. http://dx.doi.org/10.1016/j.mambio.2011.02.003.

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8

Selvan, K. Muthamizh, Gopi Govindan Veeraswami, and Syed Ainul Hussain. "Dietary preference of the Asiatic wild dog (Cuon alpinus)." Mammalian Biology 78, no. 6 (November 2013): 486–89. http://dx.doi.org/10.1016/j.mambio.2013.08.007.

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9

IYENGAR, A., V. N. BABU, S. HEDGES, A. B. VENKATARAMAN, N. MACLEAN, and P. A. MORIN. "Phylogeography, genetic structure, and diversity in the dhole (Cuon alpinus)." Molecular Ecology 14, no. 8 (July 2005): 2281–97. http://dx.doi.org/10.1111/j.1365-294x.2005.02582.x.

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10

Paulraj, S., N. Sundararajan, A. Manimozhi, and Sally Walker. "Reproduction of the Indian wild dog (Cuon alpinus) in captivity." Zoo Biology 11, no. 4 (1992): 235–41. http://dx.doi.org/10.1002/zoo.1430110403.

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11

Taron, Ulrike H., Johanna L. A. Paijmans, Axel Barlow, Michaela Preick, Arati Iyengar, Virgil Drăgușin, Ștefan Vasile, Adrian Marciszak, Martina Roblíčková, and Michael Hofreiter. "Ancient DNA from the Asiatic Wild Dog (Cuon alpinus) from Europe." Genes 12, no. 2 (January 22, 2021): 144. http://dx.doi.org/10.3390/genes12020144.

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The Asiatic wild dog (Cuon alpinus), restricted today largely to South and Southeast Asia, was widespread throughout Eurasia and even reached North America during the Pleistocene. Like many other species, it suffered from a huge range loss towards the end of the Pleistocene and went extinct in most of its former distribution. The fossil record of the dhole is scattered and the identification of fossils can be complicated by an overlap in size and a high morphological similarity between dholes and other canid species. We generated almost complete mitochondrial genomes for six putative dhole fossils from Europe. By using three lines of evidence, i.e., the number of reads mapping to various canid mitochondrial genomes, the evaluation and quantification of the mapping evenness along the reference genomes and phylogenetic analysis, we were able to identify two out of six samples as dhole, whereas four samples represent wolf fossils. This highlights the contribution genetic data can make when trying to identify the species affiliation of fossil specimens. The ancient dhole sequences are highly divergent when compared to modern dhole sequences, but the scarcity of dhole data for comparison impedes a more extensive analysis.
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Riordan, Philip, Jun Wang, Kun Shi, Hongyan Fu, Zhu Dabuxilike, Kebiao Zhu, and Xiaohu Wang. "New evidence of dhole Cuon alpinus populations in north-west China." Oryx 49, no. 2 (April 2015): 203–4. http://dx.doi.org/10.1017/s0030605315000046.

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13

Venkataraman, Arun B., R. Arumugam, and R. Sukumar. "The foraging ecology of dhole (Cuon alpinus) in Mudumalai Sanctuary, southern India." Journal of Zoology 237, no. 4 (December 1995): 543–61. http://dx.doi.org/10.1111/j.1469-7998.1995.tb05014.x.

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14

Volodina, Elena V., Ilya A. Volodin, Irina V. Isaeva, and Carolyn Unck. "Biphonation May Function to Enhance Individual Recognition in the Dhole, Cuon alpinus." Ethology 112, no. 8 (August 2006): 815–25. http://dx.doi.org/10.1111/j.1439-0310.2006.01231.x.

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15

Yadav, Shailendra Kumar, Babu Ram Lamichhane, Naresh Subedi, Ramesh Kumar Thapa, Laxman Poudyal, and Bhagawan Raj Dahal. "Dhole Cuon alpinus (Mammalia: Carnivora: Canidae) rediscovered in Bardia National Park, Nepal." Journal of Threatened Taxa 11, no. 12 (September 26, 2019): 14582–86. http://dx.doi.org/10.11609/jott.4714.11.12.14582-14586.

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An increasing intensity of camera traps recorded the presence of poorly known and globally Endangered Asiatic Wild Dogs Cuon alpinus from different locations in recent years in Nepal. After 18 years since the previous report, we recorded 29 photos and a video of Dholes in four independent detections with an effort of 4,035 trap-nights during camera trap surveys targeted at tigers in the winter of 2016/2017. Solitary dholes were camera-trapped from four locations within 27.45km2 area in Bardia National Park. The evidence of a dead Dhole probably killed in retaliation shows the threat to the species. Dholes co-exist in Bardia with sympatric carnivores like Tiger Panthera tigris, Leopard Panthera pardus, and Jackal Canis aureus.
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16

MAISCH, H. "The influence of husbandry and pack management on Dhole Cuon alpinus reproduction." International Zoo Yearbook 44, no. 1 (January 2010): 149–64. http://dx.doi.org/10.1111/j.1748-1090.2009.00090.x.

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17

Aryal, Achyut, Saroj Panthi, Rosemary K. Barraclough, Roberta Bencini, Bikash Adhikari, Weihong Ji, and David Raubenheimer. "Habitat selection and feeding ecology of dhole (Cuon alpinus) in the Himalayas." Journal of Mammalogy 96, no. 1 (February 15, 2015): 47–53. http://dx.doi.org/10.1093/jmammal/gyu001.

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18

Pal, Ranjana, Shagun Thakur, Shashank Arya, Tapajit Bhattacharya, and Sambandam Sathyakumar. "Recent records of dhole (Cuon alpinus, Pallas 1811) in Uttarakhand, Western Himalaya, India." Mammalia 82, no. 6 (November 27, 2018): 614–17. http://dx.doi.org/10.1515/mammalia-2017-0017.

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Abstract Camera trap photographs of solitary individuals of Asiatic wild dog or dholes (Cuon alpinus, Pallas 1811) have been recorded from Kheda Tal area in Uttarkashi district of Uttarakhand. This is the first photographic confirmation of the presence of this species in the Himalayan habitats of Uttarakhand. The presence of dholes here seems to be confined to a relatively small area of unprotected habitats affected by various anthropogenic uses. The population of endangered dhole is rapidly declining especially in Himalayan region. Thus, it is imperative that local stakeholders are aware of their presence, and it is highly recommended that investigations of the ecological parameters of this population are continued.
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19

Kazi, A. A., D. N. Rabari, M. I. Dahya, and S. Lyngdoh. "Reappearance of Dhole Cuon alpinus (Mammalia: Carnivora: Canidae) in Gujarat after 70 years." Journal of Threatened Taxa 13, no. 6 (May 26, 2021): 18655–59. http://dx.doi.org/10.11609/jott.6415.13.6.18655-18659.

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The Dhole Cuon alpinus used to be the meso carnivore of the forests throughout Indian subcontinent; however, habitat loss, low prey biomass, and human disturbance exterminated the species from India’s 60% historic range, and the numbers are less than 1,500 individuals in wild. Following the same shrinking trend, Dholes were extirpated from Gujarat. A few doubtful sightings and inevident reportings generated ambiguity of Dhole presence in Gujarat. We conducted a study in Vansda National Park with 15,660 trap nights at 30 trap locations, and have confirmed the rediscovery of Dholes in Gujarat after 70 years. We estimated the Dhole’s minimum home range as 13.7km2 and also analyzed relative abundance index of other mammals. The future retention of Dholes requires detailed range, diet, and adaption studies along with conservational efforts to reduce re-extinction probabilities.
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Punjabi, Girish A., Linnea Worsøe Havmøller, Rasmus Worsøe Havmøller, Dusit Ngoprasert, and Arjun Srivathsa. "Methodological approaches for estimating populations of the endangered dhole Cuon alpinus." PeerJ 10 (February 22, 2022): e12905. http://dx.doi.org/10.7717/peerj.12905.

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Large carnivores are important for maintaining ecosystem integrity and attract much research and conservation interest. For most carnivore species, estimating population density or abundance is challenging because they do not have unique markings for individual identification. This hinders status assessments for many threatened species, and calls for testing new methodological approaches. We examined past efforts to assess the population status of the endangered dhole (Cuon alpinus), and explored the application of a suite of recently developed models for estimating their populations using camera-trap data from India’s Western Ghats. We compared the performance of Site-Based Abundance (SBA), Space-to-Event (STE), and Time-to-Event (TTE) models against current knowledge of their population size in the area. We also applied two of these models (TTE and STE) to the co-occurring leopard (Panthera pardus), for which density estimates were available from Spatially Explicit Capture–Recapture (SECR) models, so as to simultaneously validate the accuracy of estimates for one marked and one unmarked species. Our review of literature (n = 38) showed that most assessments of dhole populations involved crude indices (relative abundance index; RAI) or estimates of occupancy and area of suitable habitat; very few studies attempted to estimate populations. Based on empirical data from our field surveys, the TTE and SBA models overestimated dhole population size beyond ecologically plausible limits, but the STE model produced reliable estimates for both the species. Our findings suggest that it is difficult to estimate population sizes of unmarked species when model assumptions are not fully met and data are sparse, which are commonplace for most ecological surveys in the tropics. Based on our assessment, we propose that practitioners who have access to photo-encounter data on dholes across Asia test old and new analytical approaches to increase the overall knowledge-base on the species, and contribute towards conservation monitoring of this endangered carnivore.
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Gimranov, D. O., P. A. Kosintsev, and N. G. Smirnov. "The late Neopleistocene dhole (Carnivora, Canidae, Cuon alpinus Pallas, 1811) from the Urals." Doklady Biological Sciences 464, no. 1 (September 2015): 248–50. http://dx.doi.org/10.1134/s0012496615050117.

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Srivathsa, Arjun, Sushma Sharma, Priya Singh, Girish A. Punjabi, and Madan K. Oli. "A strategic road map for conserving the Endangered dhole Cuon alpinus in India." Mammal Review 50, no. 4 (July 29, 2020): 399–412. http://dx.doi.org/10.1111/mam.12209.

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Kamler, Jan F., Arlyne Johnson, Chanthavy Vongkhamheng, and Anita Bousa. "The diet, prey selection, and activity of dholes (Cuon alpinus) in northern Laos." Journal of Mammalogy 93, no. 3 (June 28, 2012): 627–33. http://dx.doi.org/10.1644/11-mamm-a-241.1.

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Frey, Roland, Ilya A. Volodin, Guido Fritsch, and Elena V. Volodina. "Potential Sources of High Frequency and Biphonic Vocalization in the Dhole (Cuon alpinus)." PLOS ONE 11, no. 1 (January 5, 2016): e0146330. http://dx.doi.org/10.1371/journal.pone.0146330.

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Dar, Shahid A., and Jamal A. Khan. "Food Habits of Dhole Cuon alpinus in Tropical Forests of Southern India." Current Science 111, no. 10 (November 1, 2016): 1701. http://dx.doi.org/10.18520/cs/v111/i10/1701-1705.

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George, Samuel, Ashokkumar Mohanarangan, Dencin Rons Thampy, and George Chandy. "Prey Composition of Post-denning Dhole (Cuon alpinus) in South Wayanad Forest Division, Kerala, India." Proceedings of the Zoological Society 74, no. 2 (March 30, 2021): 147–54. http://dx.doi.org/10.1007/s12595-020-00357-z.

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Palei, Nimain Charan, Bhakta Padarbinda Rath, Sanath Kumar, and Himanshu Shekhar Palei. "Occurrence and Activity Pattern of Endangered Dhole (Cuon alpinus) in Debrigarh Wildlife Sanctuary, Odisha, India." Proceedings of the Zoological Society 75, no. 1 (November 30, 2021): 134–38. http://dx.doi.org/10.1007/s12595-021-00391-5.

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Jenks, Kate E., Nucharin Songsasen, Budsabong Kanchanasaka, Peter Leimgruber, and Todd K. Fuller. "Local People's Attitudes and Perceptions of Dholes (Cuon Alpinus) around Protected Areas in Southeastern Thailand." Tropical Conservation Science 7, no. 4 (December 2014): 765–80. http://dx.doi.org/10.1177/194008291400700413.

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Hayward, M. W., S. Lyngdoh, and B. Habib. "Diet and prey preferences of dholes (Cuon alpinus): dietary competition within Asia's apex predator guild." Journal of Zoology 294, no. 4 (September 18, 2014): 255–66. http://dx.doi.org/10.1111/jzo.12171.

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Jog, Maithili M., Rahul R. Marathe, Shantanu S. Goel, Sachin P. Ranade, Krushnamegh K. Kunte, and Milind G. Watve. "Sarcocystis infection in Chital (Axis axis) and Dhole (Cuon alpinus) in two Indian protected areas." Zoos' Print Journal 18, no. 10 (September 21, 2003): 1220–22. http://dx.doi.org/10.11609/jott.zpj.18.10.1220-2.

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Zakir, Tania, Harish Debbarma, and Muntasir Akash. "Dhole Cuon alpinus in Satchari National Park: on the first verifiable evidence from northeast Bangladesh." Mammalia 84, no. 6 (November 26, 2020): 587–93. http://dx.doi.org/10.1515/mammalia-2019-0050.

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AbstractThe distribution range of the globally Endangered dhole Cuon alpinus overlaps mixed evergreen mid-hill forests of northeastern Bangladesh but lacks any authentic documentation. With the first evidence from this region, we observed the activity pattern of dhole, of its prey and of human activities in Satchari National Park, a part of a larger northeastern forested area, Raghunandan Hill Reserve Forest – the northernmost fringe of the Baramura Hills of India. We obtained 32 photos of solitary individuals on eight different events from six out of 587 camera trap days. Naïve occupancy estimate for dhole was 0.41 with a detection probability of 0.65. We identified two individuals, and reported five predation events. We encountered a moderately high temporal overlap of dhole activity with that of barking deer (∆̂1 = 0.65, CI = 0.42–0.83), human activity (0.63, 0.38–0.89), and livestock movement (0.59, 0.38–0.89). Given that dhole is a diurnal hyper-carnivore, illegal logging and tourism pressure appeared as major disturbances to the species' existence in the park. Habitats of northeastern protected forests, their adjacency with hill ranges of Tripura, and dhole being a habitat generalist, we conjecture its presence in the entire region and recommend thorough camera-trapping efforts.
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Thing, Santa Bahadur, Jhamak Bahadur Karki, Babu Ram Lamichhane, Shashi Shrestha, Uba Raj Regmi, and Rishi Ranabhat. "Distribution and habitat-use of Dhole Cuon alpinus (Mammalia: Carnivora: Canidae) in Parsa National Park, Nepal." Journal of Threatened Taxa 14, no. 3 (March 26, 2022): 20703–12. http://dx.doi.org/10.11609/jott.6178.14.3.20703-20712.

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Dhole Cuon alpinus is one of the top predators in Asian forests but is one of the least studied species of carnivores. We surveyed an area of 499 km2 of Parsa National Park (PNP) during the winter (November–January) of 2016–17 using camera-traps to determine the spatial distribution and habitat-use patterns of Dhole. We overlaid 2 x 2 km2 grid cells (n= 126) across the study area and set up a pair of motion sensor cameras in each grid cells for 21 days. We modeled the habitat-use by Dholes as a function of sampling covariates and fine-scale habitat covariates using single species single season occupancy models. We estimated the parameters in two steps. First, we defined a global model for probability of habitat-use and modeled detection probability (p) either as an intercept-only model or as a function of covariates. Second, we modeled the habitat-use probability (Ψ) incorporating the top-ranked model for probability of detection (p) in the first step. A total effort of 2,520 camera-trap-nights resulted in 63 independent detections of dholes at 27 locations in PNP. The naïve occupancy estimate of Dholes in PNP was 0.21. The estimated probability of habitat-use (Ψ) and detection (p) were 0.47±0.27 and 0.24±0.05, respectively. Grassland availability (βG= 8.00±3.09), terrain ruggedness index (βTRI= 0.73±0.34), and Sambar (prey) presence (βS= 1.06±0.51) strong positive association, whereas, stream/exposed surfaces (βSES= -0.45±0.43) had negative association with the habitat use by Dholes. Similarly, detection probability was positively associated with presence of Sambar (βS= 2.44±1.02) but negatively associated with streams/exposed surfaces (βSES= -0.99±0.32) and terrain ruggedness (βTRI= -0.09±0.23). Our study provides quantitative information on the ecology of Dholes with potential applications for improving their conservation efforts in Nepal.
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Venkataraman, Arun B. "Male-biased Adult Sex Ratios and their Significance for Cooperative Breeding in Dhole, Cuon alpinus, Packs." Ethology 104, no. 8 (April 26, 2010): 671–84. http://dx.doi.org/10.1111/j.1439-0310.1998.tb00101.x.

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Lyngdoh, Salvador, Govindan Veeraswami Gopi, K. Muthamizh Selvan, and Bilal Habib. "Effect of interactions among ethnic communities, livestock and wild dogs (Cuon alpinus) in Arunachal Pradesh, India." European Journal of Wildlife Research 60, no. 5 (September 4, 2014): 771–80. http://dx.doi.org/10.1007/s10344-014-0846-8.

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Ngoprasert, Dusit, George A. Gale, and Andrew J. Tyre. "Abundance estimation from multiple data types for group-living animals: An example using dhole (Cuon alpinus)." Global Ecology and Conservation 20 (October 2019): e00792. http://dx.doi.org/10.1016/j.gecco.2019.e00792.

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Zhang, Honghai, and Lei Chen. "The complete mitochondrial genome of dhole Cuon alpinus: phylogenetic analysis and dating evolutionary divergence within canidae." Molecular Biology Reports 38, no. 3 (September 22, 2010): 1651–60. http://dx.doi.org/10.1007/s11033-010-0276-y.

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Smith, Bradley P., Teghan A. Lucas, Rachel M. Norris, and Maciej Henneberg. "Brain size/body weight in the dingo (Canis dingo): comparisons with domestic and wild canids." Australian Journal of Zoology 65, no. 5 (2017): 292. http://dx.doi.org/10.1071/zo17040.

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Endocranial volume was measured in a large sample (n = 128) of free-ranging dingoes (Canis dingo) where body size was known. The brain/body size relationship in the dingoes was compared with populations of wild (Family Canidae) and domestic canids (Canis familiaris). Despite a great deal of variation among wild and domestic canids, the brain/body size of dingoes forms a tight cluster within the variation of domestic dogs. Like dogs, free-ranging dingoes have paedomorphic crania; however, dingoes have a larger brain and are more encephalised than most domestic breeds of dog. The dingo’s brain/body size relationship was similar to those of other mesopredators (medium-sized predators that typically prey on smaller animals), including the dhole (Cuon alpinus) and the coyote (Canis latrans). These findings have implications for the antiquity and classification of the dingo, as well as the impact of feralisation on brain size. At the same time, it highlights the difficulty in using brain/body size to distinguish wild and domestic canids.
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Singh, Priya, Arjun Srivathsa, and David W. Macdonald. "Conservation status of the dhole Cuon alpinus in north-east India, with a focus on Dampa Tiger Reserve, Mizoram." Oryx 54, no. 6 (October 23, 2019): 873–77. http://dx.doi.org/10.1017/s0030605319000255.

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AbstractDespite the efforts invested in their conservation, the status of many threatened carnivores in key conservation landscapes remains unknown. The dhole Cuon alpinus is an Endangered social carnivore whose geographical range has contracted by c. 80% since the early 20th century. North-east India is a critical link between South Asian and South-east Asian dhole populations. In this study we compiled presence records of dholes across north-east India, from multiple sources. We also conducted camera-trap surveys in one part of this region, Dampa Tiger Reserve in the state of Mizoram. We examined the influence of ecological and management factors on fine-scale site-use by dholes in Dampa Tiger Reserve, showing a positive association of dhole site-use with sambar Rusa unicolor encounters, distance to the forest boundary and presence of forest department personnel, underscoring the importance of prey and protection. Our findings also highlight the need for targeted, multi-scale assessments of dhole ecology across other sites in north-east India.
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Nurvianto, Sandy, Rury Eprilurahman, Muhammad Ali Imron, and Sven Herzog. "FEEDING HABITS OF PACK LIVING DHOLE (Cuon alpinus) IN A DRY DECIDUOUS FOREST OF EAST JAVA, INDONESIA." Taprobanica 8, no. 1 (March 26, 2016): 10–20. http://dx.doi.org/10.47605/tapro.v8i1.210.

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Malez, Mirko, and Ivan Turk. "Cuon alpinus europaeus Bourguignat (Carnivora, Mammalia) from the Upper Pleistocene in the Cave Apnarjeva jama at Celje." Geologija 33, no. 1 (June 30, 1990): 215–32. http://dx.doi.org/10.5474/geologija.1990.004.

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41

Xue, Yadong, Diqiang Li, Wenfa Xiao, Yuguang Zhang, Bin Feng, and Heng Jia. "Records of the dhole (Cuon alpinus) in an arid region of the Altun Mountains in western China." European Journal of Wildlife Research 61, no. 6 (July 24, 2015): 903–7. http://dx.doi.org/10.1007/s10344-015-0947-z.

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42

Wirsing, Aaron J., and Thomas M. Newsome. "Scavenging Effects of Large Canids." Integrative and Comparative Biology 61, no. 1 (March 21, 2021): 117–31. http://dx.doi.org/10.1093/icb/icab012.

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Synopsis Many large predators are also facultative scavengers that may compete with and depredate other species at carcasses. Yet, the ecological impacts of facultative scavenging by large predators, or their “scavenging effects,” still receive relatively little attention in comparison to their predation effects. To address this knowledge gap, we comprehensively examine the roles played by, and impacts of, facultative scavengers, with a focus on large canids: the African wild dog (Lycaon pictus), dhole (Cuon alpinus), dingo (Canis dingo), Ethiopian wolf (Canis simensis), gray wolf (Canis lupus), maned wolf (Chrysocyon brachyurus), and red wolf (Canis rufus). Specifically, after defining facultative scavenging as use or usurpation of a carcass that a consumer has not killed, we (1) provide a conceptual overview of the community interactions around carcasses that can be initiated by facultative scavengers, (2) review the extent of scavenging by and the evidence for scavenging effects of large canids, (3) discuss external factors that may diminish or enhance the effects of large canids as scavengers, and (4) identify aspects of this phenomenon that require additional research attention as a guide for future work.
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Charaspet, Khwanrutai, Ronglarp Sukmasuang, Noraset Khiowsree, Nucharin Songsasen, Saksit Simchareon, and Prateep Duengkae. "Some ecological aspects of dhole (Cuon alpinus) in the Huai Kha Khaeng Wildlife Sanctuary, Uthai Thani Province, Thailand." Folia Oecologica 46, no. 2 (December 1, 2019): 91–100. http://dx.doi.org/10.2478/foecol-2019-0012.

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AbstractThe dhole (Cuon alpinus) is one of the least frequent studied endangered canid species and many aspects of ecological knowledge about this species are lacking. The objectives of this study were to investigate the spatial movement of dholes, prey abundance, prey selection, and prey overlaps with other large carnivorous species in the Huai Kha Khaeng Wildlife Sanctuary, Thailand, during November, 2017 and October, 2018. Two adult female dholes were captured and fitted with GPS collars. Twenty camera trap sets were systematically used to survey the area. Scat collection was conducted along forest roads and trails. The home range sizes and activity radii of the two dholes were 3,151.63 ha. and 1,442.84 m, and 33.39 ha and 331.56 m, respectively. The sambar deer (Rusa unicolor) was the most abundant prey species (30.93%). However, dhole fecal analysis showed that the monitored dholes preferred red muntjac (Muntiacus muntjak) (57.1%). There was a high degree of prey overlap between dholes and leopards (98%), indicating very high prey competition. The dholes in this study represent movement patterns in richly abundant prey habitats, but with the presence of other predators that can affect prey selection and movement patterns of the dhole in the area.
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Karanth, K. Ullas, and Melvin E. Sunquist. "Behavioural correlates of predation by tiger (Panthera tigris), leopard (Panthera pardus) and dhole (Cuon alpinus) in Nagarahole, India." Journal of Zoology 250, no. 2 (February 2000): 255–65. http://dx.doi.org/10.1111/j.1469-7998.2000.tb01076.x.

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45

Ghezzo, Elena, and Lorenzo Rook. "Cuon alpinus (Pallas, 1811) (Mammalia, Carnivora) from Equi (Late Pleistocene, Massa-Carrara, Italy): anatomical analysis and palaeoethological contextualisation." Rendiconti Lincei 25, no. 4 (November 8, 2014): 491–504. http://dx.doi.org/10.1007/s12210-014-0345-6.

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46

Jain, Manjari, Singha Utpal, and S. Mukhopadhyay. "Indirect Evidences of Wildlife Activities in Shoals of Western Ghats, a Biodiversity Hotspots." Vestnik Zoologii 45, no. 2 (January 1, 2011): e-33-e-39. http://dx.doi.org/10.2478/v10058-011-0010-9.

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Indirect Evidences of Wildlife Activities in Shoals of Western Ghats, a Biodiversity HotspotsThe presence of wildlife fauna and its activities were ascertained with the density of the scat, dung and other markings or droppings of the wildlife abode therein. Attempt was made to find out spatial differences in the activities of the wildlife populations and to comment on the abundance of different preys and predators within shola forests of Western Ghat hill forests, a Biodiversity hotspot in India. An indirect sampling method, Transect Count Method, was employed to count dung/pellet group/scat and other markings in that area. Pachyderms were found to be mostly dominant in Varagaliar and Punnumala shola patches while scats of all the three important predators, viz., tiger (Panthera tigris tigris), Indian wild dog (Cuon alpinus) and leopard (Panthera pardus) were encountered only in Varagaliar shola. Greater abundance was recorded from Indira Gandhi Wildlife Sanctuary than Silent Valley National Park may be because of the restriction of animal movements in the former due to topographical barriers and its existence as isolated shola patches that led to a greater concentration of wild fauna in a relatively segregated forest cover.
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Karanth, K. Ullas, Arjun Srivathsa, Divya Vasudev, Mahi Puri, Ravishankar Parameshwaran, and N. Samba Kumar. "Spatio-temporal interactions facilitate large carnivore sympatry across a resource gradient." Proceedings of the Royal Society B: Biological Sciences 284, no. 1848 (February 8, 2017): 20161860. http://dx.doi.org/10.1098/rspb.2016.1860.

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Species within a guild vary their use of time, space and resources, thereby enabling sympatry. As intra-guild competition intensifies, such behavioural adaptations may become prominent. We assessed mechanisms of facilitating sympatry among dhole ( Cuon alpinus ), leopard ( Panthera pardus ) and tiger ( Panthera tigris ) in tropical forests of India using camera-trap surveys. We examined population-level temporal, spatial and spatio-temporal segregation among them across four reserves representing a gradient of carnivore and prey densities. Temporal and spatial overlaps were higher at lower prey densities. Combined spatio-temporal overlap was minimal, possibly due to chance. We found fine-scale avoidance behaviours at one high-density reserve. Our results suggest that: (i) patterns of spatial, temporal and spatio-temporal segregation in sympatric carnivores do not necessarily mirror each other; (ii) carnivores are likely to adopt temporal, spatial, and spatio-temporal segregation as alternative mechanisms to facilitate sympatry; and (iii) carnivores show adaptability across a gradient of resource availability, a driver of inter-species competition. We discuss behavioural mechanisms that permit carnivores to co-occupy rather than dominate functional niches, and adaptations to varying intensities of competition that are likely to shape structure and dynamics of carnivore guilds.
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Wolf, Christopher, and William J. Ripple. "Prey depletion as a threat to the world's large carnivores." Royal Society Open Science 3, no. 8 (August 2016): 160252. http://dx.doi.org/10.1098/rsos.160252.

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Large terrestrial carnivores are an ecologically important, charismatic and highly endangered group of species. Here, we assess the importance of prey depletion as a driver of large carnivore endangerment globally using lists of prey species for each large carnivore compiled from the literature. We consider spatial variation in prey endangerment, changes in endangerment over time and the causes of prey depletion, finding considerable evidence that loss of prey base is a major and wide-ranging threat among large carnivore species. In particular, the clouded leopard ( Neofelis nebulosa ), Sunda clouded leopard ( Neofelis diardi ), tiger ( Panthera tigris ), dhole ( Cuon alpinus ) and Ethiopian wolf ( Canis simensis ) all have at least 40% of their prey classified as threatened on the International Union for the Conservation of Nature (IUCN) Red List and, along with the leopard ( Panethra pardus ), all of these species except the Ethiopian wolf have at least 50% of their prey classified as declining. Of the 494 prey species in our analysis, an average of just 6.9% of their ranges overlap protected areas. Together these results show the importance of a holistic approach to conservation that involves protecting both large carnivores directly and the prey upon which they depend.
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Modi, Shrushti, Bilal Habib, Pallavi Ghaskadbi, Parag Nigam, and Samrat Mondol. "Standardization and validation of a panel of cross-species microsatellites to individually identify the Asiatic wild dog (Cuon alpinus)." PeerJ 7 (September 2, 2019): e7453. http://dx.doi.org/10.7717/peerj.7453.

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Background The Asiatic wild dog or dhole (Cuon alpinus) is a highly elusive, monophyletic, forest dwelling, social canid distributed across south and Southeast Asia. Severe pressures from habitat loss, prey depletion, disease, human persecution and interspecific competition resulted in global population decline in dholes. Despite a declining population trend, detailed information on population size, ecology, demography and genetics is lacking. Generating reliable information at landscape level for dholes is challenging due to their secretive behaviour and monomorphic physical features. Recent advances in non-invasive DNA-based tools can be used to monitor populations and individuals across large landscapes. In this paper, we describe standardization and validation of faecal DNA-based methods for individual identification of dholes. We tested this method on 249 field-collected dhole faeces from five protected areas of the central Indian landscape in the state of Maharashtra, India. Results We tested a total of 18 cross-species markers and developed a panel of 12 markers for unambiguous individual identification of dholes. This marker panel identified 101 unique individuals from faecal samples collected across our pilot field study area. These loci showed varied level of amplification success (57–88%), polymorphism (3–9 alleles), heterozygosity (0.23–0.63) and produced a cumulative misidentification rate or PID(unbiased) and PID(sibs) value of 4.7 × 10−10 and 1.5 × 10−4, respectively, indicating a high statistical power in individual discrimination from poor quality samples. Conclusion Our results demonstrated that the selected panel of 12 microsatellite loci can conclusively identify dholes from poor quality, non-invasive biological samples and help in exploring various population parameters. This genetic approach would be useful in dhole population estimation across its range and will help in assessing population trends and other genetic parameters for this elusive, social carnivore.
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Grassman, Lon I., Jan E. Janecka, Sean C. Austin, Michael E. Tewes, and Nova J. Silvy. "Chemical immobilization of free-ranging dhole (Cuon alpinus), binturong (Arctictis binturong), and yellow-throated marten (Martes flavigula) in Thailand." European Journal of Wildlife Research 52, no. 4 (April 11, 2006): 297–300. http://dx.doi.org/10.1007/s10344-006-0040-8.

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