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1

Singh, Surbhi, Geeta Maheshwari, and Insha Nihal. "Taxonomic Morphology of Biting Midges (Diptera: Ceratopogonidae)." UTTAR PRADESH JOURNAL OF ZOOLOGY 44, no. 22 (November 6, 2023): 244–53. http://dx.doi.org/10.56557/upjoz/2023/v44i223739.

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Ceratopogononidae is a family of small nematocerous flies ranging approximately 1 to 3mm in length. Ceratopogonidae which was previously categorised as a subfamily of Chironomidae is distinguised by the presence of its biting mouthparts. Ceratopogonids commonly known as biting midges are the vectors of numerous pathogens such as viruses, protozoans and nematodes. Hence its transmission has a great veterinary and medical importance. This family is divided into four subfamilies: Leptoconopinae, Forcipomyiinae, Dasyhilinae and Ceratopogoninae. The flies were collected with the help of aspirator, Diptera net and UV light trap. The collected specimens were preserved in the 70% ethanol and later dissected out with the help of Digital Motic Dissecting Binocular. Ceratopogonidae include widely diversified morphotaxanomic character describing various subgroups. This variable character which include colour pattern, sizes of the body, proportions of various morphological parts are not very reliable for taxonomic identification. The study found that the morphotaxanomic feature like external genitalia of male and female, armature and arrangement of wing venation are much more stable. There are several puzzling problems in the determination of disputed species, can be resolved on the basis of taxonomic characters of male and female genitalia. The present paper will resolve the problem related to plethora of taxonomic character with special reference to biting midges.
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DA SILVA, IRENE MARQUES, RUTH L. FERREIRA-KEPPLER, NEUSA HAMADA, and CARLA G. CAZORLA. "Redescription of Stilobezzia (Stilobezzia) albicoxa Lane & Forattini, 1956 with new synonymy (Diptera: Ceratopogonidae)." Zootaxa 4822, no. 3 (August 6, 2020): 443–45. http://dx.doi.org/10.11646/zootaxa.4822.3.10.

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Stilobezzia Kieffer, 1911 is a diverse genus of predaceous Ceratopogonidae (Ceratopogoninae: Ceratopogonini) with 349 species recognized worldwide (Borkent & Dominiak 2020). The genus has four subgenera, three occurs in Brazil, where Stilobezzia s. str. is the most speciose, with 23 species registered (Santarém & Felippe-Bauer 2019).
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3

Szadziewski, Ryszard. "Biting midges (Diptera: Ceratopogonidae) as indicators of biostratigraphy, ecological reconstructions and identification of amber deposits." Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107, no. 2-3 (June 2016): 219–30. http://dx.doi.org/10.1017/s1755691017000378.

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ABSTRACTBiting midges (Diptera: Ceratopogonidae) are a large family of flies that commononly appear in Lower Cretaceous to Miocene strata, with over 280 fossil species (4.3 % of the family), belonging to 49 genera (26 extant; 23 extinct). Morphological characters used in the identification of fossil genera and species are identical to those used in studies of extant Ceratopogonidae and, as a result, their potential indicative value is reliable. Two relictual extant genera, Leptoconops and Austroconops, reported from Lower Cretaceous Lebanese amber, are at least 125 million years old. Certain ceratopogonid genera are indicative for the Lower Cretaceous, Upper Cretaceous, Eocene or Miocene. A morphological character indicative for the Upper Cretaceous and Cenozoic is macrotrichia on the wing membrane of adults. Indicator species and genera are reviewed for all amber deposits. Eocene Baltic amber contains the best known fauna of biting midges, with 109 named species. Some genera are indicative of aquatic and semiaquatic habitats (predaceous genera, subfamily Ceratopogoninae), forests with rotting trees (Forcipomyia), sandy sea shore habitats (Leptoconops), a cold boreal climate (Ceratopogon) or warm climates (Nannohelea, Austroconops, Leptoconops, Meunierohelea, Metahelea). Females require a protein-rich meal and are well known for feeding on the blood of vertebrates, but many feed on other things, so this information can help with the interpretation of palaeoenvironments. Washingtonhelea taimyrica Szadziewski, 1996, described from Siberian amber, is transferred to the fossil genus Palaeobrachypogon: P.taimyricus (Szadziewski, 1996), comb. nov. For Serromyia alphea, mistakenly redescribed and illustrated from Eocene Bitterfeld amber (= Baltic amber) by Szadziewski (1993), a new name – Serromyiaerrata Szadziewski, nom. nov. – is proposed.
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4

Tóthová, Andrea, Jan Knoz, Miroslav Barták, and Štěpán Kubík. "Biomonitoring of Ceratopogonidae (Diptera: Nematocera) using car nets." Entomologica Fennica 16, no. 2 (June 1, 2005): 124–28. http://dx.doi.org/10.33338/ef.84245.

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Car nets were used to collect two samples on a forest road in Podyji National park, the Braitava forest, Czech Republic. Sampling was done in 2002 between May 31 and June 1, and between July 30 and 31. These consisted of 10 rounds (each 10 km in length) from morning to dusk. Over 3,000 specimens (52 species) of Ceratopogonidae were captured. The results suggest that the car-net method may be efficient in ceratopogonid biomonitoring and e. g. determining their daily flight activity and swarming sites.
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5

Golovatyuk, Larisa V., Tatiana D. Zinchenko, Nadezhda N. Sushchik, Galina S. Kalachova, and Michail I. Gladyshev. "Biological aspects of the associations of biting midges (Diptera: Ceratopogonidae) in two saline rivers of the Elton Lake Basin, Russia." Marine and Freshwater Research 69, no. 6 (2018): 906. http://dx.doi.org/10.1071/mf17125.

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We studied species composition, density, biomass and production of larvae of the family Ceratopogonidae in two saline rivers (Volgograd region, Russia). Ceratopoponids make up an important part of macroinvertebrate community in these rivers. Average monthly production (dry weight) of ceratopogonid larvae in the rivers was 3.5–4.8gm–2 month–1 in May and ~0.9gm–2 month–1 in August. For the first time, feeding spectra of ceratopogonid larvae, Palpomyia schmidti Goetghebuer, 1934, was studied using fatty acid analyses. The larvae of P. schmidti appeared to selectively consume diatoms and other algae and to avoid bacteria and decomposed dead organic matter (detritus) of low nutritive quality.
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6

Carlos Henrique Marchiori. "Biology and feeding behavior of ceratopogonid adult (Diptera: Ceratopogonidae)." International Journal of Frontiers in Science and Technology Research 1, no. 2 (December 30, 2021): 007–24. http://dx.doi.org/10.53294/ijfstr.2021.1.2.0073.

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The purpose of this paper is to understand the biology and feeding behavior of ceratopogonid adult (Diptera: Ceratopogonidae). The research was carried out in studies related to quantitative aspects taxonomic and conceptual aspects such as habitat, geographical distribution, traps for collecting, their parasitoids and predator, life cycle, damage, economic importance, medicinal importance, biological aspects, and reproduction. A literature search was carried out containing articles published from 1993 to 2021. The mini review was prepared in Goiânia, Goiás, from September to October 2021, through the. The mini-review was prepared in Goiânia, Goiás, from September to October 2021, through the Online Scientific Library (Scielo), internet, ResearchGate, Academia.edu, Frontiers, Publons, Qeios, Portal of Scientific Journals in Health Sciences, Pubmed, Online Scientific Library (Scielo), internet, ResearchGate, Academia.edu, Frontiers, Biological Abstract, Publons, Qeios, Portal of Scientific Journals in Health Sciences, and Pubmed, Dialnet, World, Wide Science, Springer, RefSeek, Microsoft Academic, Science, ERIC, Science Research.com, SEEK education, Periódicos CAPES, Google Academic, Bioline International, VADLO, Scopus, and Web of Science.
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7

SZADZIEWSKI, RYSZARD, and MARTA GWIZDALSKA-KENTZER. "New records of biting midges (Diptera: Ceratopogonidae) from the United Arab Emirates, with a description of a new species." Zootaxa 4894, no. 4 (December 11, 2020): 594–97. http://dx.doi.org/10.11646/zootaxa.4894.4.6.

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Biting midges (Ceratopogonidae) are a relatively well-studied family of nematocerous flies (Borkent & Dominiak 2020), but the ceratopogonid fauna of the Middle East is not well known. To date, only 264 species from this family have been recorded in this entire region (Alwin et al. 2016a, b) and just 53 in the United Arab Emirates (UAE) (Szadziewski et al. 2011, Alwin et al. 2016a, b). Here, we report on three species of biting midges new to the fauna of the UAE.
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8

Metz, Robert. "Sinusoidal trail formed by a Recent biting midge (Family Ceratopogonidae): trace fossil implications." Journal of Paleontology 61, no. 2 (March 1987): 312–14. http://dx.doi.org/10.1017/s0022336000028481.

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A Recent sinusoidal trail made by a biting midge (family Ceratopogonidae, genus ?Bezzia) moving on the surface of wet mud along the edge of a freshwater pond has been observed. The trail is similar to the trace fossil Cochlichnus Hitchcock, 1858. The wave length, wave amplitude, and overall form and shape of this modern trail compares favorably to Eocene wave-like trails attributed to nematodes. Consequently, a ceratopogonid larva is believed responsible for at least some of those fossil trails.
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9

Borkent, Art. "The Ceratopogonidae (Diptera) of the Galápagos Islands, Ecuador with a discussion of their phylogenetic relationships and zoogeographic origins." Insect Systematics & Evolution 22, no. 1 (1991): 97–122. http://dx.doi.org/10.1163/187631291x00336.

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AbstractEleven species of Ceratopogonidae are recognized on the Galápagos Islands. Four of these appear to be endemic, six are widespread occurring elsewhere on the continental Americas and one is of uncertain distributional status. One of the endemic species is described as new, Dasyhelea sinclairi Borkent. Each of the species is most closely related to taxa occurring elsewhere than on the Galápagos Islands (with one possible but unlikely exception of two of these being sister species), indicating at least ten independent invasions of the islands by the ancestors or ancestral populations of these species. It is uncertain whether the widespread species have been brought through the agency of man or through natural dispersal to the Galápagos Islands. Based on comparisons with the Ceratopogonidae fauna of oceanic islands elsewhere, the presence of endemic species only in the genera Forcipomyia and Dasyhelea on the Galápagos Islands is likely due to the superior dispersal capabilities of members of these genera. This may also explain the presence of those ceratopogonid species occurring both on the Galápagos Islands and the New World mainland.
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10

Perkovsky, E. E. "Comparison of Biting Midges of the Early Eocene Cambay Amber (India) and Late Eocene European Ambers Supports the Independent Origin of European Ambers." Vestnik Zoologii 51, no. 4 (August 28, 2017): 275–84. http://dx.doi.org/10.1515/vzoo-2017-0033.

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Abstract Of the three genera of Ceratopogoninae dominant in Late Eocene ambers of Europe, the two known are absent in the Early Eocene Cambay amber (India): the Holarctic genus Ceratopogon and the worldwide distributed genus Culicoides, known since the Late Cretaceous. Tropical biting midge genera (Meunierohelea and Leptoconops and genera with feminized male antennae: Camptopterohelea, Eohelea, and Gedanohelea) are abundantly represented in Cambay amber. The proportion of Ceratopogonini with feminized male antennae among all Ceratopogonidae dramatically increases from north to south: from 1.4-2.4 % in unbiased collections of Bitterfeld amber (Humboldt Museum collection) and Baltic amber (the Giecewicz collection) to 7 % in Danish amber and 12.7 % in Rovno amber; their proportion in Cambay amber is 17 %. The proportion of tropical specimens among specimens Ceratopogoninae in unbiased collections is 6.4 % in Baltic amber, 5.3 % in Bitterfeld amber, 21 % in Rovno and Danish amber, and 58 % in Cambay amber. Strong differences in the proportion of tropical components among Ceratopogoninae from different European ambers are in agreement with data on Cambay amber and so are indicative of origin of the European ambers under different climatic and hence geographic environments.
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11

Macfie, J. W. S. "NOTES ON CERATOPOGONIDAE." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 18, no. 5-6 (March 18, 2009): 109–15. http://dx.doi.org/10.1111/j.1365-3113.1949.tb01431.x.

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12

Arnold, Sarah E. J., Samantha J. Forbes, David R. Hall, Dudley I. Farman, Puran Bridgemohan, Gustavo R. Spinelli, Daniel P. Bray, et al. "Floral Odors and the Interaction between Pollinating Ceratopogonid Midges and Cacao." Journal of Chemical Ecology 45, no. 10 (October 2019): 869–78. http://dx.doi.org/10.1007/s10886-019-01118-9.

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AbstractMost plant species depend upon insect pollination services, including many cash and subsistence crops. Plants compete to attract those insects using visual cues and floral odor which pollinators associate with a reward. The cacao tree, Theobroma cacao, has a highly specialized floral morphology permitting pollination primarily by Ceratopogonid midges. However, these insects do not depend upon cacao flowers for their life cycle, and can use other sugar sources. To understand how floral cues mediate pollination in cacao we developed a method for rearing Ceratopogonidae through several complete lifecycles to provide material for bioassays. We carried out collection and analysis of cacao floral volatiles, and identified a bouquet made up exclusively of saturated and unsaturated, straight-chain hydrocarbons, which is unusual among floral odors. The most abundant components were tridecane, pentadecane, (Z)-7-pentadecene and (Z)-8-heptadecene with a heptadecadiene and heptadecatriene as minor components. We presented adult midges, Forcipomyia sp. (subgen. Forcipomyia), Culicoides paraensis and Dasyhelea borgmeieri, with natural and synthetic cacao flower odors in choice assays. Midges showed weak attraction to the complete natural floral odor in the assay, with no significant evidence of interspecific differences. This suggests that cacao floral volatiles play a role in pollinator behavior. Midges were not attracted to a synthetic blend of the above four major components of cacao flower odor, indicating that a more complete blend is required for attraction. Our findings indicate that cacao pollination is likely facilitated by the volatile blend released by flowers, and that the system involves a generalized odor response common to different species of Ceratopogonidae.
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13

Borkent, Art, Robert A. Coram, and Edmund A. Jarzembowski. "The oldest fossil biting midge (Diptera: Ceratopogonidae) from the Purbeck Limestone Group (Lower Cretaceous) of southern Great Britain." Polish Journal of Entomology / Polskie Pismo Entomologiczne 82, no. 4 (December 1, 2013): 273–79. http://dx.doi.org/10.2478/v10200-012-0041-8.

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ABSTRACT The oldest fossil Ceratopogonidae, Archiaustroconops besti sp. n., dated at 142 million years old, is described from the Purbeck Limestone Group of Dorset, southern England, United Kingdom. Represented by a single wing, it belongs to a lineage within the family, indicating that the fossil subfamily Lebanoculicoidinae, the earliest lineage, may be expected in even earlier deposits. The wing, like those of all early fossil Ceratopogonidae is small, less than 1 mm in length.
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14

Macfie, J. W. S. "NOTES ON CERATOPOGONIDAE (DIPTERA)." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 6, no. 6 (March 18, 2009): 111–18. http://dx.doi.org/10.1111/j.1365-3113.1937.tb00451.x.

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15

Macfie, J. W. S. "NOTES ON CERATOPOGONIDAE (DIPTERA)." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 7, no. 8 (March 18, 2009): 157–66. http://dx.doi.org/10.1111/j.1365-3113.1938.tb01271.x.

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Macfie, J. W. S. "NOTES ON CERATOPOGONIDAE (DIPTERA)." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 10, no. 5 (March 18, 2009): 67–69. http://dx.doi.org/10.1111/j.1365-3113.1941.tb00696.x.

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Macfie, J. W. S. "CERATOPOGONIDAE (DIPTERA) FROM EGYPT." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 12, no. 9-10 (March 18, 2009): 145–59. http://dx.doi.org/10.1111/j.1365-3113.1943.tb00763.x.

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18

SPINELLI, GUSTAVO A., and MARTA WOLFF. "FAMILY CERATOPOGONIDAE." Zootaxa 4122, no. 1 (June 14, 2016): 98. http://dx.doi.org/10.11646/zootaxa.4122.1.15.

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19

Clastrier, J. "Ceratopogonidae de Nouvelle-Calédonie." Annales de Parasitologie Humaine et Comparée 60, no. 6 (1985): 747–59. http://dx.doi.org/10.1051/parasite/1985606747.

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20

Sontag, Elżbieta, and Ryszard Szadziewski. "Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine)." Polish Journal of Entomology / Polskie Pismo Entomologiczne 80, no. 4 (December 1, 2011): 779–800. http://dx.doi.org/10.2478/v10200-011-0058-4.

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Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine) The paper presents the results of an examination of 714 biting midges (Diptera: Ceratopogonidae) preserved in Baltic amber from the Rovno deposits in Ukraine. A new species - Leptoconops rovnensis sp. n. - is described and illustrated. 29 of the fossil species reported here have already been described from other deposits of Baltic amber: 26 of these were also found in amber from the Gulf of Gdańsk and 18 in amber from Bitterfeld (Saxony). The most common genera of biting midges in Ukrainian amber are also found in amber from Bitterfeld and the Gulf of Gdańsk, and with very much the same frequencies. The results indicate that the faunas of Ceratopogonidae enclosed in amber from Rovno, Bitterfeld and the Baltic are very similar, showing that they inhabited similar palaeoenvironments in the same palaeogeographic region.
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21

Lewis, David J. "BITING FLIES (DIPTERA) OF PEATLANDS AND MARSHES IN CANADA." Memoirs of the Entomological Society of Canada 119, S140 (1987): 133–40. http://dx.doi.org/10.4039/entm119140133-1.

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AbstractA review of the literature on biting flies of Canadian peatlands and marshes reveals that 21 species of Culicidae, 43 species of Tabanidae, and 6 species of Ceratopogonidae are known to occur in such habitats. Of these 70 species of biting flies, 45 occur in bogs, 22 in fens, and 36 in marshes. Although the species composition of wetland culicids is largely known, immatures of many species of the tabanids and ceratopogonids have yet to be collected and/or described. Almost nothing is known about the biology of the immature stages in wetland habitats. Most species known to inhabit wetlands have been recorded from both peatlands and marshes, but there are representatives of all families that appear to be restricted to a particular type of wetland. Compounding this paucity of information on the biting flies of Canadian wetlands is the inconsistency and lack of specificity in describing wetland habitats.
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22

MEILLON, BOTHA. "NOTES ON AFRICAN CERATOPOGONIDAE (DIPTERA)." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 7, no. 12 (March 18, 2009): 266–70. http://dx.doi.org/10.1111/j.1365-3113.1938.tb01237.x.

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23

Kremer, M., and E. Lienhart. "Élevage deCulicoides nubeculosus(Diptera: Ceratopogonidae)." Parasite 5, no. 3 (September 1998): 211–14. http://dx.doi.org/10.1051/parasite/1998053211.

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24

Direni Mancini, José Manuel, Cecilia Adriana Veggiani-Aybar, Ana Denise Fuenzalida, Mercedes Sara Lizarralde de Grosso, and María Gabriela Quintana. "Ceratopogonidae (Diptera: Nematocera) of the piedmont of the Yungas forests of Tucumán: ecology and distribution." PeerJ 4 (November 17, 2016): e2655. http://dx.doi.org/10.7717/peerj.2655.

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Within the Ceratopogonidae family, many genera transmit numerous diseases to humans and animals, while others are important pollinators of tropical crops. In the Yungas ecoregion of Argentina, previous systematic and ecological research on Ceratopogonidae focused on Culicoides, since they are the main transmitters of mansonelliasis in northwestern Argentina; however, few studies included the genera Forcipomyia, Dasyhelea, Atrichopogon, Alluaudomyia, Echinohelea, and Bezzia. Therefore, the objective of this study was to determine the presence and abundance of Ceratopogonidae in this region, their association with meteorological variables, and their variation in areas disturbed by human activity. Monthly collection of specimens was performed from July 2008 to July 2009 using CDC miniature light traps deployed for two consecutive days. A total of 360 specimens were collected, being the most abundant Dasyhelea genus (48.06%) followed by Forcipomyia (26.94%) and Atrichopogon (13.61%). Bivariate analyses showed significant differences in the abundance of the genera at different sampling sites and climatic conditions, with the summer season and El Corralito site showing the greatest abundance of specimens. Accumulated rainfall was the variable that related the most to the abundance of Culicoides (10.56%), while temperature was the most closely related variable to the abundance of Forcipomyia, Dasyhelea, and Atrichopogon.
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25

Borkent, Art. "The Lower Cretaceous male ofLebanoculicoides daheri– belonging to the earliest lineage of biting midges (Diptera: Ceratopogonidae)." Canadian Entomologist 151, no. 3 (March 14, 2019): 278–90. http://dx.doi.org/10.4039/tce.2019.4.

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AbstractRe-examination of the male biting midge ofLebanoculicoides daheriChoufani, Azar, and Nel (Diptera: Ceratopogonidae), in 127 million-year-old Lebanese amber, revealed further features and details of morphological, phylogenetic, and bionomic importance. The phylogenetic position of the fossil genusLebanoculicoidesSzadziewski as the sister group of all remaining extant and extinct Ceratopogonidae is confirmed. A revised key to all four known species of this genus is provided. A permanently erect antennal plume is hypothesised as an additional synapomorphy ofAustroconopsWirth and Lee and two Cretaceous fossil genera,MinyoheleaBorkent andArchiaustroconopsSzadziewski. The presence of a hind tibial comb and more distal row of spines is considered a synapomorphy of all Culicomorpha other than Chironomidae, with some secondary losses within this group.
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Felippe-Bauer, Maria L., and Cristiane S. Silva. "Morphological alterations in Neotropical Ceratopogonidae (Diptera)." Revista Brasileira de Zoologia 23, no. 3 (September 2006): 593–96. http://dx.doi.org/10.1590/s0101-81752006000300001.

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Morphological alterations in six different species of females Culicoides Latreille, 1809 and one of Monohelea Kieffer, 1917 from Brazil, Mexico, Panama and Peru are described. The correlation of the morphological changes with the taxonomy and behavior of the species is discussed.
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27

Macfie., J. W. S. "CERATOPOGONIDAE (DIPT.) FROM THE RIVER AMAZON." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 4, no. 3 (March 18, 2009): 49–56. http://dx.doi.org/10.1111/j.1365-3113.1935.tb00555.x.

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28

Macfie, J. W. S. "CERATOPOGONIDAE FROM THE ANGLO-EGYPTIAN SUDAN." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 16, no. 5-6 (March 18, 2009): 69–78. http://dx.doi.org/10.1111/j.1365-3113.1947.tb00861.x.

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29

Hasan;YAĞCI, EREN. "ANKARA'DA BULUNAN CULİCOİDES (DİPTERA: CERATOPOGONİDAE) TÜRLERİ." Ankara Üniversitesi Veteriner Fakültesi Dergisi 42, no. 2 (1995): 1. http://dx.doi.org/10.1501/vetfak_0000000776.

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30

Spinelli, Gustavo R. "Synonymy in Neotropical Ceratopogonidae (Diptera: Culicomorpha)." Revista de la Sociedad Entomológica Argentina 78, no. 3 (September 26, 2019): 38–39. http://dx.doi.org/10.25085/rsea.780308.

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31

Meillon, Botha. "CERATOPOGONIDAE (DIPTERA, NEMATOCERA) FROM SOUTHERN AFRICA." Transactions of the Royal Entomological Society of London 90, no. 16 (April 24, 2009): 455–66. http://dx.doi.org/10.1111/j.1365-2311.1940.tb01030.x.

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32

Vorsprach, Bettina, Christian Karl Meiser, Doreen Werner, Carsten Balczun, and Günter A. Schaub. "Monitoring of Ceratopogonidae in Southwest Germany." Parasitology Research 105, no. 2 (April 2, 2009): 337–44. http://dx.doi.org/10.1007/s00436-009-1411-3.

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33

Barceló, Carlos, and Ignacio Ruiz-Arrondo. "Diversity, Distribution and Phylogeny of Vector Insects." Diversity 16, no. 2 (February 1, 2024): 95. http://dx.doi.org/10.3390/d16020095.

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Mosquitoes (Fam. Culicidae), sand flies (Subfam. Phlebotominae), biting midges(Fam. Ceratopogonidae), black flies (Fam. Simuliidae) and stable flies (Fam. Muscidae) are groups of insects capable to transmit pathogens of public health and veterinary importance [...]
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34

Stuart, A. E., A. Evans, C. Brooks, M. J. A. Simpson, J. B. Cloughley, D. F. MacIntosh, C. L. E. Stuart, A. Blackwell, and D. S. Kettle. "The Biting Midge of the West Highlands: Fifty Years of Research." Scottish Medical Journal 41, no. 5 (October 1996): 143–46. http://dx.doi.org/10.1177/003693309604100505.

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The biting midge of the West Highlands belongs to the family Ceratopogonidae and approximately 150 species are known to exist in Britain. All of the flies are of minute size and slender build with wings which fold over the back. The females have biting mouthparts including a needle sharp proboscis with scissor type mandibles. The males do not bite. The Ceratopogonidae devours other small insects, some feed on plant juices, others pierce the wing veins of butterflies and some attack juicy caterpillars. Only three genera are bloodsuckers and of these only Culicoides occurs in Britain. The members of this genus are classified by the pattern of wing venation and the spots on the wings. Edwards' gives a detailed table of these characteristics. The commonest species in Scotland is C. impunctatus, although others have been met with on Skye.
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35

Golovatyuk, L. V. "Salinity Tolerance, Seasonal and Multiyear Dynamics of Biting Midges (Diptera, Ceratopogonidae) in Macrozoobenthos Communities of Saline Rivers (the Lake Elton Basin, Russia)." Биология внутренних вод, no. 6 (November 1, 2023): 842–52. http://dx.doi.org/10.31857/s0320965223060128.

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The results of long-term studies of biting midges (Diptera, Ceratopogonidae) in macrozoobenthos communities of five saline rivers of the hyperhaline Lake Elton basin are presented. The fauna of Ceratopogonidae includes 5 halophilic and euryhaline species, among which representatives of the tribe Palpomyiinae predominate. Taxonomic composition of the family differs in rivers with different levels of salinity. Species Sphaeromias miricornis and Culicoides riethi are widely distributed in mesohaline the Khara, the Lantsug and the Bolshaya Samoroda rivers, and P. schmidti widely distributed in polyhaline the Chernavka and the Solyanka rivers. Two generations of S. miricornis and three generations of P. schmidti were recorded. Departure of the first generation of P. schmidti occurs in mid-May, S. miricornis in late May–early June. There is an increase in mean annual abundance and biomass of biting midges in polyhaline rivers compared to mesohaline rivers.
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36

TÓTHOVÁ, ANDREA, and PABLO I. MARINO. "A new Nearctic species of Atrichopogon (Meleohelea) and a redescription of Atrichopogon (M.) chilensis Ingram & Macfie (Diptera: Ceratopogonidae)." Zootaxa 2023, no. 1 (February 27, 2009): 47–54. http://dx.doi.org/10.11646/zootaxa.2023.1.3.

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A new species of Ceratopogonidae from Canada, Atrichopogon (Meloehelea) ladislavi Tóthová, is described. The Patagonian species Atrichopogon (M.) chilensis Ingram & Macfie is restored from synonymy, its female is redescribed and the male is described for the first time.
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37

Grogan, William L., Lawrence J. Hribar, and Francis G. Howarth. "The Old World biting midge, Forcipomyia (Lepidohelea) pulcherrima SANTOS ABREU, new to the fauna of the United States (Diptera: Ceratopogonidae)." Polish Journal of Entomology / Polskie Pismo Entomologiczne 82, no. 4 (December 1, 2013): 287–302. http://dx.doi.org/10.2478/v10200-012-0043-6.

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ABSTRACT We provide the first United States records of the Old World biting midge, Forcipomyia (Lepidohelea) pulcherrima SANTOS ABREU (Diptera: Ceratopogonidae), from California, Florida and Hawaii. The fourth instar larva of F. pulcherrima is also described and illustrated for the first time.
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38

Clastrier, Jean, and Jean-Claude Delécolle. "Ceratopogonidae des îles Wallis et Futuna (Diptera)." Bulletin de la Société entomologique de France 101, no. 3 (1996): 289–318. http://dx.doi.org/10.3406/bsef.1996.17256.

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39

Spinelli, Gustavo R., and Willis W. Wirth. "The Neotropical Species of Phaenobezzia (Diptera: Ceratopogonidae)." Florida Entomologist 69, no. 1 (March 1986): 231. http://dx.doi.org/10.2307/3494765.

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40

Macfie, J. W. S. "CERATOPOGONIDAE (DIPTERA) FROM ETHIOPIA AND BRITISH SOMALILAND." Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 6, no. 4 (March 18, 2009): 73–79. http://dx.doi.org/10.1111/j.1365-3113.1937.tb00301.x.

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41

Weerekoon, A. C. J. "ON THE BEHAVIOUR OF CERTAIN CERATOPOGONIDAE (DIPTERA)." Proceedings of the Royal Entomological Society of London. Series A, General Entomology 28, no. 7-9 (April 2, 2009): 85–92. http://dx.doi.org/10.1111/j.1365-3032.1953.tb00649.x.

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42

Glick, Jayson I. "Culicoides Biting Midges (Diptera: Ceratopogonidae) of Kenya." Journal of Medical Entomology 27, no. 2 (March 1, 1990): 85–195. http://dx.doi.org/10.1093/jmedent/27.2.85.

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43

Holbrook, Frederick R., and Walter J. Tabachnick. "Culicoides variipennis (Diptera: Ceratopogonidae) Complex in California." Journal of Medical Entomology 32, no. 4 (July 1, 1995): 413–19. http://dx.doi.org/10.1093/jmedent/32.4.413.

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44

Mair, J., and A. Blackwell. "Mating Behavior of Culicoides nubeculosus (Diptera: Ceratopogonidae)." Journal of Medical Entomology 33, no. 5 (September 1, 1996): 856–58. http://dx.doi.org/10.1093/jmedent/33.5.856.

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45

Nunamaker, R. A., S. E. Brown, and D. L. Knudson. "Metaphase Chromosomes of Culicoides variipennis (Diptera: Ceratopogonidae)." Journal of Medical Entomology 33, no. 5 (September 1, 1996): 871–73. http://dx.doi.org/10.1093/jmedent/33.5.871.

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46

Schmidtmann, E. T., F. R. Holbrook, E. Day, T. Taylor, and W. J. Tabachnick. "Culicoides variipennis (Diptera: Ceratopogonidae) Complex in Virginia." Journal of Medical Entomology 35, no. 5 (September 1, 1998): 818–24. http://dx.doi.org/10.1093/jmedent/35.5.818.

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47

Braverman, Y., A. Chizov-Ginzburg, and B. A. Mullens. "Mosquito Repellent Attracts Culicoides imicola (Diptera: Ceratopogonidae)." Journal of Medical Entomology 36, no. 1 (January 1, 1999): 113–15. http://dx.doi.org/10.1093/jmedent/36.1.113.

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48

Spinelli, Gustavo R. "Erratum: Synonymy in Neotropical Ceratopogonidae (Diptera: Culicomorpha)." Revista de la Sociedad Entomológica Argentina 79, no. 2 (June 29, 2020): 1. http://dx.doi.org/10.25085/rsea.790201.

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LINLEY, J. R., and G. M. ADAMS. "Sexual receptivity in Culicoides melleus (Diptera: Ceratopogonidae)." Transactions of the Royal Entomological Society of London 126, no. 3 (April 24, 2009): 279–303. http://dx.doi.org/10.1111/j.1365-2311.1974.tb00855.x.

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50

Debenham, M. L. "CERATOPOGONIDAE (DIPTERA) OF THE COCOS (KEELING) ISLANDS." Australian Journal of Entomology 26, no. 4 (November 1987): 347–48. http://dx.doi.org/10.1111/j.1440-6055.1987.tb01980.x.

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