Dissertations / Theses on the topic 'Broken River'

In lowland river systems, in-channel, slow-flowing or still-water areas (still-water patches, SWPs) are considered to be important habitats for many organisms, particularly the early-life stages of fish and shrimp. However, the distribution of the early life-stages of fish and shrimp among these habitats appears to be very patchy and studies suggest that the quality and diversity of microhabitat conditions within SWPs and the accessibility of SWPs to spawning adults and dispersing young may be important determinants of their suitability as nursery habitat. The aims of this thesis were to examine the utilisation of still-water patches by fish and shrimp in a lowland river in relation to habitat suitability and accessibility, with particular emphasis on early-life stages. To determine the factors influencing habitat selection among SWPs, the environmental variability in SWP habitat, and both the distribution and the movement patterns of fish and shrimp, were examined in the Broken River, a lowland river in south-eastern Australia. SWP habitat was found to be highly spatially and temporally variable in the Broken River. SWPs differed in relation to permanence, accessibility and microhabitat variables, and all life-stages of fish and shrimp were found to be significantly spatially aggregated among SWPs. This suggests that individual SWPs may differ in their suitability as habitat, and/or in their accessibility to dispersing organisms and indicates either differential rates of retention, movement into SWPs, spawning effort or survival among SWPs for these organisms. Significant associations were found for all species and life-stages in relation to the microhabitat characteristics of SWPs. The two introduced species, carp and gambusia, were found to have fewer associations, which suggests that these species are habitat generalists. Cover and SWP morphology variables were shown to be important for all native species. Significant, positive associations were found for most species and life-stages with large, deep, SWPs containing instream cover, however, the extent of cover preferred was variable. It was hypothesised that large, deep SWPs that contain instream cover are more environmentally stable and provide better foraging efficiency and reduced competition for space, whilst also providing refuge from predators and, that they may be easier to locate than smaller patches. Specific associations with microhabitat variables differed among all species and life-stages, and this was attributed to differences in diet and predation rates. Consequently, generalised microhabitat relationships for particular life-stages or species could not be identified and the results from this thesis suggest that a diversity of microhabitat conditions are required to meet the differing requirements of various life-stages and species. Significant associations were also found for most groups in relation to the accessibility characteristics of SWPs, indicating that the ability of individuals to access SWPs is an important factor in determining their distribution among SWPs. This further suggests that movement is an important factor in the distribution pattern of fish and shrimp among SWPs. Significant associations were found for most groups in relation to patch isolation, adjacent hydraulic habitat and entrance conditions, indicating that landscape composition and configuration as well as boundary conditions may be important determinants of organisms being able to locate suitable patches. Associations with accessibility variables differed among species and life-stages, and may be attributable to differences in movement capabilities. Field manipulations of instream cover and entrance depth were conducted to further examine the habitat associations found. The results confirmed a positive relationship between instream cover and fish and shrimp abundances. No species, however, responded consistently to the manipulation of entrance depths, and this was attributed to water level rises throughout the experiment and/or the correlation of entrance depth with SWP depth. However, the results from the field manipulations suggested that deeper habitats are able to be exploited by small-bodied adults and larvae when significant levels of instream cover are also available as refuge from predation. In order to confirm the importance of movement in the selection of SWP habitat by fish and shrimp, the movement patterns of fish and shrimp into and out of SWPs were investigated. Whilst the results from this aspect of the study were inconclusive for fish, the results for shrimp confirmed that adults and larvae moved routinely into and out of SWPs. However, for all shrimp species, movement appeared to be limited to a certain period of larval development, indicating that SWP quality and stability may be more important at particular stages of development than others. The results of this thesis have demonstrated the importance of SWP quality and stability for fish and shrimp in the Broken River and have shown that habitat preferences vary among individual species and life-stages. Consequently, in order to manage for multiple species and life-stages, consideration must be given not only to the availability of SWPs, but also to their stability over time and to the availability of a diverse range of microhabitats. In addition, consideration must also be given to the accessibility of SWPs and this will require a greater knowledge of the specific spawning and dispersal requirements of the organisms which utilise these patches, in combination with a greater understanding of the impacts of flow modification on riverine landscape composition and configuration.

Detailed provenance analysis and glauconite morphology of the Broken River and Iron CreeklWaipara Formations and other correlatives were conducted at 6 localities: the MandamusDove River area, Waipara River, Avoca-Iron Creek, Castle Hill Basin, Malvern and Mt Somers. The basal Broken River Formation is a fluvial boulder conglomerate interbedded with sandstones, mudstones and coal dated as Haumurian (Late Cretaceous) by pollen. The transgression is marked by a gradual drowning of the fluvial conglomerates with minor glauconite appearing in the beds immediately overlying the conglomerates in alllocali ties. The estuarine-marginal marine I lower shoreface succession of the Broken River Formation contains minor nascent micaceous glauconite. This increases in the overlying Iron Creek I Waipara Formations to 30-60% glauconite composed of nascent-micaceous to evolved/mature glauconity types characteristic oflower shoreface I foreshore to shallow shelf depositional setting. Up section, evolved mature glaucony dominates, in some beds formed in situ (autochthonous) and in other beds transported from nearby (parautochthonous) to line foresets. An extremely low sedimentation rate is necessary to form the evolved I mature type of glauconite. The age of the greensands is Teurian to Whaingaroan (Late Paleocene-Late Eocene). Overall the glauconite analysis indicates extremely low sedimentation rates with autochthonous I parautochthonous glaucony formation in nearshore marine settings, possibly even estuary environments. Clast counts from the basal conglomerates indicate derivation from local sources such as the underlying Torlesse greywackes (Pahau and Rakaia Terranes) and/or the Mandamus Igneous Complex. Sandstone composition indicates the addition of more distal sources. Quartzose sandstones plot in the interior craton province in QFL plots for both Broken River and Iron Creek/Waipara Greensand Formations. Sandstone lithics are probably derived from the underlying Torlesse greywacke. Alkali feldspar dominates over plagioclase indicating a probable plutonic felsic source. SEM-cathodoluminescence on quartz grains indicates a bimodal metamorphic to plutonic quartz grains with minor volcanic input. Plutonic grains are identified by healed microcracks, and are possibly derived from Western Province plutonic suites such as the Karamea and Separation Point Batholiths. Polycrystalline/dark CL quartz grains indicate a relatively high grade metamorphic source such as the OtagolHaast Schist while dark CL monocrystalline quartz grains indicate a low to medium metamorphic grade source such as the Alpine Schist. Volcanic quartz is zoned with straight extinction and was most likely derived from the Cretaceous Mount Somers Volcanics Group. Overall the provenance suggests local derivation of sediments when coarse fluvial deposition occurred followed by more distal derivation once transported in the nearshore marine setting.

Analysis of fossils from cave deposits at Mount Etna (eastern-central Queensland) has established that a species-rich rainforest palaeoenvironment existed in that area during the middle Pleistocene. This unexpected finding has implications for several fields (e.g., biogeography/phylogeography of rainforest-adapted taxa, and the impact of climate change on rainforest communities), but it was unknown whether the Mount Etna sites represented a small refugial patch of rainforest or was more widespread. In this study numerous bone deposits in caves in north-east Queensland are analysed to reconstruct the environmental history of the area during the late Quaternary. Study sites are in the Chillagoe/Mitchell Palmer and Broken River/Christmas Creek areas. The cave fossil records in these study areas are compared with dated (middle Pleistocene-Holocene) cave sites in the Mount Etna area. Substantial taxonomic work on the Mount Etna faunas (particularly dasyurid marsupials and murine rodents) is also presented as a prerequisite for meaningful comparison with the study sites further north. Middle Pleistocene sites at Mount Etna contain species indicative of a rainforest palaeoenvironment. Small mammal assemblages in the Mount Etna rainforest sites (>500-280 ka) are unexpectedly diverse and composed almost entirely of new species. Included in the rainforest assemblages are lineages with no extant representatives in rainforest (e.g., Leggadina), one genus previously known only from New Guinea (Abeomelomys), and forms that appear to bridge gaps between related but morphologically-divergent extant taxa ('B-rat' and 'Pseudomys C'). Curiously, some taxa (e.g., Melomys spp.) are notable for their absence from the Mount Etna rainforest sites. After 280 ka the rainforest faunas are replaced by species adapted to open, dry habitats. At that time the extinct ‘rainforest’ dasyurids and rodents are replaced by species that are either extant or recently extant. By the late Pleistocene all ‘rainforest’ and several ‘dry’ taxa are locally or completely extinct, and the small mammal fauna resembles that found in the area today. The faunal/environmental changes recorded in the Mount Etna sites were interpreted by previous workers as the result of shifts in climate during the Pleistocene. Many samples from caves in the Chillagoe/Mitchell-Palmer and Broken River/Christmas Creek areas are held in the Queensland Museum’s collection. These, supplemented with additional samples collected in the field as well as samples supplied by other workers, were systematically and palaeoecologically analysed for the first time. Palaeoecological interpretation of the faunal assemblages in the sites suggests that they encompass a similar array of palaeoenvironments as the Mount Etna sites. ‘Rainforest’ sites at the Broken River are here interpreted as being of similar age to those at Mount Etna, suggesting the possibility of extensive rainforest coverage in eastern tropical Queensland during part of the Pleistocene. Likewise, faunas suggesting open, dry palaeoenvironments are found at Chillagoe, the Broken River and Mount Etna, and may be of similar age. The 'dry' faunal assemblage at Mount Etna (Elephant hole Cave) dates to 205-170 ka. Dating of one of the Chillagoe sites (QML1067) produced a maximum age for the deposit of approximately 200 ka, and the site is interpreted as being close to that age, supporting the interpretation of roughly contemporaneous deposition at Mount Etna and Chillagoe. Finally, study sites interpreted as being of late Pleistocene-Holocene age show faunal similarities to sites of that age near Mount Etna. This study has several important implications for the biogeography and phylogeography of murine rodents, and represents a major advance in the study of the Australian murine fossil record. Likewise the survey of the northern study areas is the first systematic analysis of multiple sites in those areas, and is thus a major contribution to knowledge of tropical Australian faunas during the Quaternary. This analysis suggests that climatic changes during the Pleistocene affected a large area of eastern tropical Queensland in similar ways. Further fieldwork and dating is required to properly analyse the geographical extent and timing of faunal change in eastern tropical Queensland.

The aim of this study was to investigate population densities and dynamics of brown trout along multiple tributaries of the Taieri River, a large New Zealand river. Relationships between juvenile brown trout population dynamics and food availability, discharge and water temperature were examined in the same lowland stream. Downstream movements of juvenile brown trout were also examined. Trace element analyses of eggs aimed to identify the source of fish spawning in tributary streams. An experimental study of juvenile brown trout collected from downstream and upstream reaches compared patterns of agonistic behaviour amongst trout that were considered to be dominated by either migratory (downstream) or resident (upstream) fish. Longitudinal surveys of juvenile trout abundance were expanded across four additional streams to determine whether there were consistent patterns in population dynamics of brown trout. Large spatial and temporal variation in population dynamics of juvenile brown trout was found in Silverstream with an upstream site having significantly less temporal variation in densities than a downstream site. No effect of food availability on population dynamics was found. Discharge and temperature affected population densities of juvenile brown trout in some years, but could not solely explain the general pattern in spatial variation in population dynamics. Downstream movement was found to be a behavioural strategy used by juvenile brown trout and appeared to contribute to variation in juvenile density. Analysis of trace element signatures in brown trout eggs indicated that upstream spawning migration by anadromous females was restricted by the presence of a steep gorge along the middle reaches of the Taieri River. Patterns of spawning by migratory and resident fish along tributary streams appeared to vary. In Silverstream, spawning by anadromous fish dominated the lower reaches, whereas considerable overlap between anadromous and resident fish was observed in Big Stream. In Cap Burn distinct trace element signatures were observed for redds spawned along upstream and downstream reaches. However, it was not possible to confidently ascribe the source of the redds to main river migrants or Cap Burn resident fish. Behavioural observations of juvenile brown trout collected from upstream (resident) and downstream (migratory) reaches of Silverstream revealed differences in aggression level, with migratory fish being more aggressive. No difference in growth rates was found between fish from the two areas. Furthermore, the social organisation of resident fish was based on the formation of dominance hierarchies, whereas the competitive status of migratory fish was more even. Patterns in spatial and temporal variation in the longitudinal population dynamics and patterns of abundance of juvenile trout were repeated across the five tributaries. Populations in upstream reaches consisted of fish from multiple age classes and exhibited relatively limited variation in densities, whereas juvenile fish dominated populations along downstream reaches and densities varied greatly over an annual cycle. Taken together, these results suggest a general pattern of spatial and temporal variation in the population dynamics of brown trout sub-populations along tributaries of a large New Zealand river. Whilst various environmental factors may contribute to some of the variation, longitudinal separation into resident and migratory populations also appeared to be important. These finding are important for the management of brown trout in both main river channels and tributaries in New Zealand.

A multi-scale approach for restoration site selection is presented and applied to an upland catchment, the River Ure, North Yorkshire. Traditional survey methods, advances in remote sensing, Geographical Information Systems (GIS) and risk-based fine sediment modelling using the SCIMAP module are combined to gather data at the catchment-scale through to the in-stream habitat-scale. The data gathered have been assessed against spatially distributed brown trout fry populations using Pearson’s correlation and multiple stepwise regressions. Fine sediment was shown to have a positive correlation with fry populations when upland drainage channels (grips) were added to the SCIMAP model. This suggests risk from peatland drainage is realised further down the catchment where eroded sediments are deposited. Farm-scale SCIMAP modelling was tested against farmers’ knowledge with variable results. It appears there is a cultural response to risk developed over generations. Management of meadows and pasture land through sub-surface drainage and stock rotation resulted in the risk being negated or re-routed across the holding. At other locations apparently low-risk zones become risky through less sensitive farming methods. This multi-scale approach reveals that the largest impacts on brown trout recruitment operate at the habitat-adjacent scale in tributaries with small upstream areas. The results show a hierarchy of impact, and risk-filters, arising from different intensity land management. This offers potential for targeted restoration site selection. In low-order streams it seems that restoration measures which exclude livestock, and provide bankside shading, can be effective. At such sites the catchment-scale shows a reduced signal on in-stream biota. Thus, brown trout stocks could be significantly enhanced by targeting restoration at riffle-habitat zones and adjacent land in order to disconnect the stream from farm-derived impacts and through adding structure to the stream channel.

Abstract Maintaining connectivity in boreal streams by rehabilitation procedures is a challenging task that requires ecological understanding based on empirical research. In this thesis, I examined the effects of stream rehabilitation on densities and growth of brown trout (Salmo trutta L.), as well as on benthic biodiversity. As streams are known to be closely connected to their watersheds, I also examined the effects of watershed drainage activities on stream organisms. The results of this thesis have implications for enhancing salmonid populations, as well for maintaining stream biodiversity. First, regional climatic variability was shown to override local impacts of watershed management on stream biota. This highlights the importance of placing results from local studies in a regional context. Second, increased sedimentation that typically follows anthropogenic actions in the watershed did not only cause direct mortality on the early life stages of brown trout, but also forced them to emerge earlier from gravel and constrained their development. Thus, sedimentation may have far-reaching fitness consequences on juvenile salmonids. Third, while instream rehabilitation did enhance habitat diversity for salmonid fish, there were only marginal effects on juvenile fish and benthic biodiversity. Therefore, the factors limiting stream biota, and obscuring positive effects of rehabilitation, are to be found elsewhere. For this purpose, my thesis offers at least three potential, not mutually exclusive explanations: (i) land use changes have altered watersheds and this can be seen as decreased stream biodiversity that cannot be corrected through local-scale restoration efforts; (ii) a habitat-forming organism group, stream bryophytes, is dispersal limited, slowing down any positive responses to restoration by fish or invertebrates that depend strongly on bryophytes; and (iii) changes to stream habitat heterogeneity caused by channelization for timber floating were rather modest to start with, and therefore any effects of stream habitat rehabilitation on stream biota are likely to be subtle. Based on these findings, I suggest that future restoration efforts should be prioritized according to a comprehensive watershed assessment. Also, monitoring of projects should be more rigorous and preferably multidisciplinary, documenting the ecological as well as hydrological and socioeconomic outcomes of rehabilitation projects.

Thesis (M.S.)--West Virginia University, 2005.
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The passage efficiency of downstream migrating brown trout (Salmo trutta) smolts past two hydropower plants in the River Emån, southeastern Sweden, was evaluated by telemetry. From 26 April to 17 May 2005, 40 wild brown trout smolts were radio-tagged, and their positions were recorded daily until 14 June. Approximately 45% of the individuals passed both power plants, and 15% made it to the sea, some 28 km downstream of the power plants. About 20% of the individuals ceased migrating, residing in a lotic habitat. Mortality due to predation was 33%, and the predation rate was higher in the power plant dams than it was downstream of the power plants. Turbine-induced mortality was greater in the upper power plant (40%) than in the lower one (13%). Fish that chose the alternative routes, through the gates, swam past the upper dam faster than fish that swam through the turbines. The results confirm previous work in the river, namely that measures need to be taken to improve passage efficiency past the power plants, particularly passage of the upper power plant. Such measures are of crucial importance if the nature-like bypass channels at the power plants in Finsjö are going to have any long-term effects on the brown trout population.

Passageeffektiviteten för nedströms vandrande havsöringssmolt (Salmo trutta) förbi två vattenkraftverk i Emån, i sydöstra Sverige, utvärderades med telemetri. Från 26 april till och med 17 maj 2005 märktes 40 vilda havsöringssmolt med radiosändare och de positionsbestämdes dagligen till och med 14 juni. Ungefär 45% av individerna passerade båda kraftverken, och 15% klarade sig till havet, cirka 28 km nedströms kraftverken. Omkring 20% av individerna avbröt vandringen och stannade i ett lotiskt habitat. Mortalitet på grund av predation uppgick till 33%, och predationshastigheten var högre i kraftverksdammarna än nedströms kraftverken. Den turbininducerade mortaliteten var högre i det övre kraftverket (40%) än i det nedre (13%). Det gick fortare att passera den övre kraftverksdammen för fisk som valde de alternativa rutterna, genom dammluckorna, jämfört med att passera genom turbinerna. Dessa resultat bekräftar tidigare studier i ån, nämligen att åtgärder krävs för att förbättra passageeffektiviteten förbi kraftverken, i synnerhet passagen av det övre kraftverket. Sådana åtgärder är av avgörande betydelse om de naturlika fiskvägarna vid kraftverken i Finsjö ska få några långsiktiga effekter på havsöringspopulationen.

Due to the significant threats posed by nonnative fish species worldwide, it is important to understand how life-history strategies of individual species interact with environmental conditions to explain the success or failure of nonnative fish invasions. Brown trout are prolific invaders, but often exhibit upstream distributional limits in Intermountain West streams, potentially due to a maladaptive reproductive life-history strategy influenced by hydrologic conditions in high-elevation areas. We used redd counts, egg survival experiments, and temperature modeling to investigate the reproductive life-history strategy of brown trout and its potential for success along an elevational stream gradient. We documented brown trout spawning in stream reaches at elevations higher than where we typically encounter brown trout during summer electrofishing surveys, indicating the potential for upstream invasion. We observed a decline in egg survival at higher elevation, cooler water sites, but did not document complete recruitment failure at these sites, again indicating the potential for successful invasion at this life-stage. Temperature data indicate that during most years, incubating brown trout eggs would likely fail to emerge from the gravel prior to peak spring flows in these high-elevation stream reaches, suggesting that damaging spring floods may cause significant egg and sac-fry mortality at high elevations, and may determine invasion success in these areas. Our results highlight the importance of identifying specific mechanisms of recruitment failure in order to better predict nonnative fish invasions in the future.

Riverine physical habitats and habitat utilization by fish have often been studied independently. Varying flows modify habitat composition and connectivity within a stream but its influence on habitat use is not well understood. This study examined brown trout (Salmo trutta) and bullhead (Cottus gobio) utilization of physical habitats that vary with flow in terms of size and type, persistence or duration, and frequency of change from one state to another, by comparing groundwater-dominated sites on the River Tern (Shropshire) with surface runoff-dominated lowland, riffle-pool sites on the Dowles Brook (Worcestershire). Mesohabitat surveys carried out at two-month intervals on a groundwater-dominated stream and on a surface runoff-influenced stream showed differences in habitat composition and diversity between the two types of rivers. The temporal variability in mesohabitat composition was also shown to differ between the two flow regime types. In the groundwater-influenced stream, mesohabitat composition hardly varied between flows whereas in the flashy stream it varied to a great extent with discharge. Habitat suitability curves for brown trout and bullhead were constructed to predict the potential location of the fish according to flow. The resulting prediction maps were tested in the field during fish surveys using direct underwater observation (snorkelling). Under the groundwater-influenced flow regime brown trout displayed a constant pattern of mesohabitat use over flows. Mesohabitats with non-varying characteristics over flows and with permanent features such as large woody debris, macrophytes or any feature providing shelter and food were favoured. Biological processes, such as hierarchy, life cycle and life stage appeared to play a key role in determining fish habitat use and to a greater extent than physical processes in these streams. Bullhead observations in the flashy river showed that mesohabitat use varied with flow but that some mesohabitats were always favoured in the stream. Pools and glides were the most commonly used mesohabitat, due to their stability over flows and their role as shelter from harsh hydraulic conditions and as food retention zones. The presence of cobbles was also found to be determinant in bullhead choice of habitat. In this flashy environment, physical processes such as flow and depth and velocity conditions appeared to be a more decisive factor in bullhead strategy of habitat use than biological processes. This research shows that: 1. Though differences in habitat use strategies between the two flow regimes can in part be attributed to differing ecology between the species, flow variability affects fish behaviour. 2. A stable flow regime allows biological processes to be the main driving force in determining fishbehaviour and location. A highly variable environment requires fish to develop behaviour strategies in response to variations in hydraulic conditions, such as depth and velocity, which constitute the key factor in determining fish location.

Abstract Despite the great amount of in-stream restorations conducted in the past decades there is still a disturbing lack of knowledge about the outcome of these measures. The overall goal of this study was to assess the effect of enhanced streambed heterogeneity on the ecology of stream salmonids and stream retention efficiency. Substratum heterogeneity is often considered as one of the most important limiting factors for organisms living in running waters. Winter ecology of rivers has not been broadly studied regardless of the general belief that wintertime conditions strongly influence the survival and population size of stream salmonids. In an experimental study, the paucity of wintertime habitat in simplified channels caused temporary mass loss in age-0 trout. In late spring, channelized stream trout performed catch-up growth with potentially negative effects on long-term fitness. A management implication of this study is that increasing cover availability by in-stream restoration structures may enhance the long term success of juvenile salmonids although the short term effects were minor. Densities of salmon parr in the River Kiiminkijoki showed no response to streambed restoration. Suitable habitat area for salmon parr increased after restoration under summer conditions. However, restoration-induced benefits to winter habitats were marginal, with one study reach indicating even negative values. Most of the areas with good habitat values were located along river margins, indicating that restoration measures had only limited impact on the mid-sections of the river channel. Dredging of small streams may have caused depletion of allochthonous organic matter due to the reduction of retentive structures. In a leaf release experiment, moss cover enhanced retentiveness as well as did various restoration structures (boulders, large wood). Only a very high amount of wood clearly enhanced retention capacity. This underlines the importance of wood as an effective retention structure in headwater streams. This study indicates that habitat complexity as such may be less important than life-stage specific habitat requirements of fish (e.g. cover for overwintering salmonids). Importantly, restoration may only be successful if the measures used target the limiting factor(s) of the ecosystem or the species; for salmonids, habitat complexity does not seem to be this factor
Tiivistelmä Uiton jälkeisten kunnostustoimenpiteiden määrä Suomessa on ollut huomattava, mutta vaikutusten arviointi, pelkästään kalastonkin kannalta, on jäänyt vähäiselle huomiolle. Tässä työssä selvitettiin kunnostusten merkitystä lohen ja taimenen poikasvaiheille, huomioiden etenkin pohjan rakenteellisen monimuotoisuuden vaikutus. Työssä selvitettiin myös kunnostusten vaikutuksia lehtikarikkeen pidätyskykyyn, joka on erityisesti latvapurojen ekosysteemien tärkeimpiä perustoimintoja. Lohikalojen talviekologinen tutkimus on viime aikoihin saakka ollut vähäistä, vaikka talviolosuhteiden uskotaan rajoittavan pohjoisten virtavesien eliöstön elinmahdollisuuksia. Kokeellisessa työssä rännimäisissä uomissa talvehtiminen aiheutti taimenenpoikasille tilapäisen painon alenemisen ja nopean kompensaatiokasvun loppukeväällä. Kompensaatiokasvu voi vaikuttaa negatiivisesti koko kalan eliniän, joten kunnostusten tuoma hyöty sopivien suojapaikkojen lisääntymisenä voi edesauttaa lohikalojen pitkäaikaista menestymistä. Kiiminkijoella lohenpoikasten tiheydet eivät muuttuneet kunnostuksen myötä ja vuosien välinen vaihtelu oli kuuden vuoden seurantajaksolla huomattavan suurta. Elinympäristömallinnuksen perusteella soveltuvan elinympäristön lisäys ei ollut merkittävää, koska etenkin talviaikaisten alueiden puute jäi huomattavaksi. Suurin osa soveltuvasta elinympäristöstä sijaitsi joen reuna-alueilla, joten kunnostusvaikutus joen keskiosaan jäi odotettua pienemmäksi. Uittoperkaus on voinut johtaa latvavesien ekosysteemien köyhtymiseen maalta tulevan orgaanisen aineksen pidättymiskyvyn vähentyessä. Kokeellisen työn perusteella kuitenkin nykypäivän tilanne vuosikymmeniä uiton loppumisen jälkeen osoittautui lähes yhtä pidättäväksi kuin nykyisin käytetyt kunnostusrakenteet (kivi tai puu). Kunnostusrakenteeseen tulisi lisätä huomattava määrä puuta, jotta lehtikarike pidättyisi korkeallakin virtaamatasolla. Tulosten perusteella elinympäristöjen muuttaminen monimuotoisemmiksi ei takaa kunnostustoimien onnistumista, sillä etenkin kalapopulaatioita rajoittavat yleensä useat tekijät. Jos kuitenkin elinympäristö on populaatiota rajoittava resurssi ja sitä pystytään lisäämään (kuten talviaikaiset suojapaikat), voidaan kunnostuksella saada näkyviä tuloksia. On ilmeistä, että kunnostustoimien tulisi olla nykyistä kattavampia ja paremmin suunnattuja rajoittaviin tekijöihin, jotta tulokset näkyisivät

Dans cette thèse, nous introduisons un nouveau modèle instationnaire de vagues valable de la zone de levée à la zone de jet de rive adapté à l'étude de la submersion. Le modèle est basé sur les équations de Serre Green-Naghdi (S-GN), dont l'application à la zone de surf reste un domaine de recherche ouvert. Nous proposons une nouvelle approche pour gérer le déferlement dans ce type de modèle, basée sur la représentation des fronts déferlés par des chocs. Cette approche a été utilisée avec succès pour les modèles basés sur les équations de Saint-Venant (SV) et permet une description simple et efficace du déferlement et des mouvements de la ligne d'eau. Dans ces travaux, nous cherchons à étendre le domaine de validité du modèle SV SURF-WB (Marche et al. 2007) vers la zone de levée en incluant les termes dispersifs propres aux équations de S-GN. Des basculements locaux vers les équations de SV au niveau des fronts permettent alors aux vagues de déferler et dissiper leur énergie. Le modèle obtenu, appelé SURF-GN, est validé à l'aide de données de laboratoire correspondant à différents types de vagues incidentes et de plages. Il est ensuite utilisé pour analyser la dynamique des fronts d'ondes longues de type tsunami en zone littorale. Nous montrons que SURF-GN peut décrire les différents types de fronts, d'ondulé non-déferlé à purement déferlé. Les conséquences de la transformation d'une onde de type tsunami en train d'ondulations lors de la propagation sur une plage sont ensuite considérées. Nous présentons finalement une étude de la célérité des vagues déferlées, basée sur les données de la campagne de mesure in-situ ECORS Truc-Vert 2008. L'influence des non-linéarités est en particulier quantifiée
In this thesis, we introduce a new numerical model able to describe wave transformation from the shoaling to the swash zones, including overtopping. This model is based on Serre Green-Naghdi equations, which are the basic fully nonlinear Boussinesq-type equations. These equations can accurately describe wave dynamics prior to breaking, but their application to the surf zone usually requires the use of complex parameterizations. We propose a new approach to describe wave breaking in S-GN models, based on the representation of breaking wave fronts as shocks. This method has been successfully applied to the Nonlinear Shallow Water (NSW) equations, and allows for an easy treatment of wave breaking and shoreline motions. However, the NSW equations can only be applied after breaking. In this thesis, we aim at extending the validity domain of the NSW model SURF-WB (Marche et al. 2007) to the shoaling zone by adding the S-GN dispersive terms to the governing equations. Local switches to NSW equations are then performed in the vicinity of the breaking fronts, allowing for the waves to break and dissipate their energy. Extensive validations using laboratory data are presented. The new model, called SURF-GN, is then applied to study tsunami-like undular bore dynamics in the nearshore. The model ability to describe bore dynamics for a large range of Froude number is first demonstrated, and the effects of the bore transformation on wave run-up over a sloping beach are considered. We finally present an in-situ study of broken wave celerity, based on the ECORS-Truc Vert 2008 field experiment. In particular, we quantify the effects of non-linearities and evaluate the predictive ability of several non-linear celerity models

On March 22, 1958 David Brower's film Wilderness Alps of Stehekin premiered to an audience of conservationists in Seattle, Washington. Almost two years in the making, the thirty-one minute film advocated the preservation of nature in Washington's North Cascades through the creation of a national park. Over the next decade, Wilderness Alps of Stehekin became the most influential publicity tool in the struggle to preserve the North Cascades. Because of the region's geographic isolation, the film was the first time many people throughout the nation were exposed to the scenic grandeur of the area. Images of craggy peaks and colorful alpine meadows resonated deeply with many Americans and persuaded them to join in the campaign. It was the voice of these citizens that led Congress to pass the North Cascades Act of 1968, which placed 674,000 acres of the North Cascades under the jurisdiction of the National Park Service. In this thesis I tell the creation story of North Cascades National Park from a conservationist perspective and trace the influence of Wilderness Alps of Stehekin within this context. Although the film was never shown in movie theaters and never aired on national television, many thousands viewed it from its premiere to the signing of the North Cascades Act. The film first introduced the idea of a North Cascades National Park, and it was important in convincing conservationists to unite around a national park solution. Ultimately, Wilderness Alps of Stehekin changed the approach activists took in the North Cascades and helped to preserve a wild and scenic nature experience for future generations through the protection of old-growth forests and alpine meadows.

Freshwater sport fisheries contribute substantially to the economies of England and Wales. However, many trout fisheries rely partly or entirely on stocking farmed trout to maintain catches within freshwater fisheries. Farmed trout often differ genetically from their wild counterparts and wild trout could be at risk of reduced fitness due to interbreeding or competition with farmed fish. Therefore, to protect remaining wild brown trout (Salmo trutta L) populations and as a conservation measure, stocking policy has changed. Legislation introduced by the Environment Agency (EA, 2009) will now only give consent to stocking of rivers and some stillwaters with sterile, all-female triploid brown trout. There are reliable triploidy induction protocols for some other commercially important salmonid species however; there is limited knowledge on triploid induction in brown trout. Previously, triploid brown trout have been produced by heat shocks although reduced survivals were obtained suggesting that an optimised heat shock had not been identified, or that heat shock gives less consistent success than hydrostatic pressure shock (HP), which is now recognised as a more reliable technique to produce triploid fish. Thus the overall aim of this thesis was to conduct novel research to support the aquaculture and freshwater fisheries sector within the United Kingdom by optimising the production and furthering the knowledge of triploid brown trout. Firstly, this PhD project investigated an optimised triploidy induction protocol using hydrostatic pressure (Chapter 2). In order to produce an optimised hydrostatic pressure induction protocol three experiments were conducted to (1) determine the optimal timing of HP shock application post-fertilisation, (2) define optimal pressure intensity and duration of the HP shock and (3) study the effect of temperature (6-12 °C) on triploid yields. Results indicated high survival to yolk sac absorption stage (69.2 - 93.6 %) and high triploid yields (82.5 - 100 %) from the range of treatments applied. Furthermore, no significant differences in triploid rates were shown when shock timings and durations were adjusted according to the temperature used. In all treatments deformity prevalence remained low during incubation (<1.8 %) up to yolk sac absorption (~550 degree days post hatch). Overall, this study indicated that the optimised pressure shock for the induction of triploidy in brown trout delivering high survival and 100 % triploid rate (a prerequisite to brown trout restocking) is a shock with a magnitude of 689 Bar applied at 300 Centigrade Temperature Minutes (CTM) for 50 CTM duration. Regarding the assessment of triploid status, the second experimental chapter tested the accuracy and efficacy of three ploidy verification techniques (Chapter 3). Techniques studied were erythrocyte nuclei measurements (Image analysis), flow cytometry (Becton Dickinson Facscalibur flow cytometer) and DNA profiling (22 polymorphic microsatellite loci) to assess the effectiveness of triploidy induction in brown trout. Results indicated the validity of using erythrocyte indices major nuclear axis measurements, flow cytometric DNA distributions expressed as relative fluorescence (FL2-Area), and polymorphic microsatellite loci (Ssa410UOS, SSa197, Str2 and SsaD48) for assessing ploidy status in brown trout. Accuracy of each technique was assessed and indicated that all techniques correctly identified ploidy level indicating 100 % triploid rate for that commercial batch of brown trout. These techniques may be utilised within aquaculture and freshwater fisheries to ensure compliance with the legislation introduced by the EA. As a result of the legislation introduced by the Environment Agency triploid brown trout will freely interact with diploid trout therefore there is a need to assess feeding response and behavioural differences between diploid and triploid trout prior to release. Therefore, in the third experimental chapter (Chapter 4) diploid and triploid brown trout were acclimated for six weeks on two feeding regimes (floating/sinking pellet). Thereafter, aggression and surface feeding response was compared between pairs of all diploid, diploid and triploid and all triploid brown trout in a semi natural stream (flume). In each pairwise matching, fish of similar size were placed in allopatry and rank determined by the total number of aggressive interactions initiated. Dominant individuals initiated more aggression than subordinates, spent more time defending a territory and positioned themselves closer to the food source (Gammarus pulex) whereas subordinates occupied the peripheries. When ploidy was considered, diploid trout were more aggressive than triploid, and dominated their siblings when placed in pairwise matchings. However, surface feeding did not differ statistically between ploidy irrespective of feeding regime. Triploids adopted a sneak feeding strategy while diploids expended more time defending a territory. In addition, an assessment of whether triploids exhibited a similar social dominance to diploids when placed in allopatry was conducted. Although aggression was lower in triploid pairs than in the diploid/triploid pairs, a dominance hierarchy was observed between individuals of the same ploidy. Dominant triploid fish were more aggressive and consumed more feed items than subordinate individuals. Subordinate fish displayed a darker colour index than dominant fish suggesting increased stress levels. However, dominant triploid fish seemed more tolerant of subordinate individuals and did not display the same degree of invasive aggression as observed in the diploid/diploid or diploid/triploid matchings. These novel findings suggest that sterile triploid brown trout feed similarly but are less aggressive than diploid trout and therefore may provide freshwater fishery managers an alternative to stocking diploid brown trout. In addition to research at the applied level in triploid brown trout, this thesis also examined the fundamental physiological effects of ploidy in response to temperature regime. Triploid salmonids have been shown to differ in their tolerance to environmental temperature. Therefore the fourth experimental chapter (Chapter 5) investigated whether temperature tolerance affected feed intake and exercise recovery. Diploid and triploid brown trout were exposed to an incremental temperature challenge (10 and 19 °C) and subsequent survival and feed intake rates were monitored. Triploids took longer to acclimate to the increase in temperature however feed intake were significantly greater in triploids at high temperature. In a follow on study, we investigated post-exercise recovery processes under each temperature regime (10 and 19 °C). Exhaustion was induced by 10 minutes of forced swimming, with subsequent haematological responses measured to determine the magnitude of recovery from exercise. Plasma parameters (alkaline phosphatase, aspartate aminotransferase, calcium, cholesterol, triglycerides, phosphorous, total protein, lactate, glucose, pH, magnesium, osmolality, potassium, sodium, chloride, lactate dehydrogenase) were measured for each ploidy. Basal samples were taken prior to exercise and then at: 1; 4, and 24 hours post-exercise. Contrary to previous studies, there was no triploid mortality during or after the exercise at either temperature. Although diploid and triploid brown trout responded metabolically to the exercise, the magnitude of the response was affected by ploidy and temperature. In particular, triploids had higher levels of plasma lactate, osmolality, and lower pH than diploids at 1 hour post exhaustive exercise. By 4 hours post-exercise plasma parameters analysed had returned to near basal levels. It was evident that the magnitude of the physiological disturbance post-exercise was greater in triploids than diploids at 19 °C. This may have implications where catch and release is practiced on freshwater fisheries. Overall, this work aimed to develop and/or refine current industry induction and assessment protocols while better understand the behaviour and physiology of diploid and triploid brown trout. The knowledge gained from this work provides aquaculture and freshwater fisheries with an optimised protocol, which delivers 100 % triploid rates and profitability without compromising farmed trout welfare, thus ultimately leading towards a more sustainable brown trout industry within the United Kingdom.