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Journal articles on the topic 'Bothremydidae'

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Published: 27 July 2024

Last updated: 29 July 2024

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1

Martín-Jiménez, Marcos, and Adán Pérez-García. "Neuroanatomical Study and Three-Dimensional Reconstruction of the Skull of a Bothremydid Turtle (Pleurodira) Based on the European Eocene Tartaruscola teodorii." Diversity 13, no. 7 (June 30, 2021): 298. http://dx.doi.org/10.3390/d13070298.

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Bothremydidae is a successful lineage of pleurodiran turtles that lived from the Cretaceous to the Paleogene, and are found in most continents of both Laurasia and Gondwana. Despite numerous known cranial remains of this clade, no study on the complete neuroanatomical reconstruction of any of its representatives has been published so far. Tartaruscola teodorii is a French lower Eocene member of Foxemydina (Bothremydini), known by two skulls. It is one of the few bothremydids identified in the Cenozoic record of Europe. The present study includes the complete three-dimensional reconstruction of each of the cranial bones of both the holotype and the paratype of T. teodorii, increasing the anatomical information about this species. The virtual reconstruction of its neuroanatomical structures is presented here, including the cranial cavity, nerves, nasal cavity, inner ears, and carotid arteries. This analysis is the first detailed neuroanatomical study performed for a member of Bothremydidae. In addition, the virtual reconstruction of the neuroanatomical structures of some extant taxa belonging to several pleurodiran lineages (Chelidae, Pelomedusidae and Podocnemididae), are also carried out and analyzed, so that the comparative framework for Pleurodira is remarkably improved.

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2

Pérez-García, Adán. "The Ornamented Shell of a New Bothremydid Turtle from the Uppermost Cretaceous of Niger." Diversity 15, no. 3 (March 5, 2023): 375. http://dx.doi.org/10.3390/d15030375.

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A new pleurodiran turtle is described here. It is identified as attributable to Bothremydidae. The new taxon comes from an upper Maastrichtian (Upper Cretaceous) outcrop located in Southwestern Niger (in the Indamane Mount area, belonging to the Abalak Department of the Tahoua Region). Abalakemys chapmanae gen. et sp. nov. is identified by an almost complete large shell of about 65 cm in length. The new bothremydid turtle is recognized as a member of Bothremydodda, showing several autapomorphies (an exclusive ornamental pattern on the plate’s outer surface, covered by small depressions; small fourth pleural scutes, only anteromedially reaching the sixth pair of costal plates; and noticeably wedged posterior plastral lobe toward the posterior region), as well as a unique combination of characters for this clade. This turtle could belong to Nigeremydini, a poorly understood Maastrichtian to Paleocene lineage of Bothremydodda, which integrates large coastal taxa that inhabited the African Trans-Saharan seaway, and for which shell information is currently extremely limited.

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3

Pereda-Suberbiola, Xabier. "Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review." Bulletin de la Société Géologique de France 180, no. 1 (January 1, 2009): 57–71. http://dx.doi.org/10.2113/gssgfbull.180.1.57.

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Abstract The continental tetrapod assemblages from the Santonian-Maastrichtian of Europe consist of dinosaurs (theropods: Abelisauroidea, Alvarezsauridae, Dromaeosauridae, ?Oviraptorosauria, ?Troodontidae, and birds: Enantiornithes, basal Ornithurae; sauropods: Titanosauria; ankylosaurs: Nodosauridae; ornithopods: Hadrosauridae, Rhabdodontidae; and neoceratopsians), pterosaurs (Azhdarchidae), crocodyliforms (eusuchians: Alligatoroidea, Gavialoidea, ?Hylaeochampsidae; sebecosuchian-like ziphosuchians; and, probably, basal neosuchians), choristoderes (?Champsosauridae), squamates (lacertilians: Iguanidae s.l., Paramacellodidae, Polyglyphanodontinae, Varanoidea; snakes: Madtsoiidae; possible amphisbaenians), turtles (cryptodires: Chelydroidea, Kallokibotionidae, Solemydidae; pleurodires: Bothremydidae, Dortokidae), lissamphibians (Albanerpetontidae; anurans: Discoglossidae, Palaeobatrachidae; caudates: Batrachosauroididae, Salamandridae), and mammals (multituberculates: Kogaionidae, ?“Paracimexomys group”; marsupials: Herpetotheriidae; eutherians: “Zhelestidae”). The palaeobiogeographical affinities of the Late Cretaceous continental tetrapods of Europe are complex. The faunas are commonly considered to show a mixed pattern resulting from the addition of “Asiamerican” and Gondwanan forms to European taxa. Albanerpetontids, both paramacellodid and polyglyphonodontine lizards, and hadrosaurid dinosaurs are taxa with Palaeolaurasian (or, in some case, even Neopangean) affinities. Other forms, such as paleobatrachid and batrachosauroidid lissamphibians, solemydid turtles, alligatoroid crocodyliforms, and nodosaurid dinosaurs can be considered as Euramerican taxa. Kallokibotionid and dortokid turtles, rhabdodontid dinosaurs and kogaionid mammals are endemic to Europe. The Gondwanan taxa have been regarded as African immigrants that reached southern Europe via the Mediterranean Tethyan sill. Abelisaurid and titanosaurid dinosaurs, trematochampsid crocodyliforms, podocnemidid and bothremydid turtles, and boid and madtsoiid snakes constitute the basic pattern of the so-called “Eurogondwanan fauna”. However, the validity of some of these taxa is a disputed matter (Titanosauridae, Trematochampsidae), and the presence of other taxa in the Late Cretaceous of Europe is based on controversial data (Boidae, Podocnemididae). Only Abelisauroidea, Madtsoiidae and Bothremydidae (and, yet for confirming, Sebecosuchia) provide evidence of interchanges between Africa and Europe. At least abelisauroids might have reached Europe before the Late Cretaceous. In conclusion, most of the continental tetrapod taxa from the latest Cretaceous of Europe show affinities with those of northern Hemisphere landmasses. Latest Cretaceous trans-Tethyan dispersal events between Africa and Europe remain poorly documented.

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4

Pérez-García, Adán. "New information on the Cenomanian bothremydid turtle <i>Algorachelus</i> based on new, well-preserved material from Spain." Fossil Record 21, no. 1 (April 13, 2018): 119–35. http://dx.doi.org/10.5194/fr-21-119-2018.

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Abstract. Algorachelus peregrinus is the oldest representative of the crown group Pleurodira known in Laurasia. The type locality of this bothremydid is Algora, situated in central Spain, at levels deposited during the uppermost middle–lowermost upper Cenomanian. A new excavation was recently carried out in this town. As a result, abundant material of Algorachelus peregrinus has been found. Several complete shells, as well as numerous partial carapaces and plastra, are presented here. The abundance of remains allows the analysis of the general patterns by which some of the shells of this littoral form were partially or totally disarticulated. The analysis of these remains not only shows several pathologies but also improves the knowledge about the anatomy and intraspecific variability of Algorachelus peregrinus. This new information allows the revision of other Cenomanian forms of Bothremydidae, both from the Middle East and from North America. Thus, Algorachelus is identified in these regions, being represented in the early or middle Cenomanian of Palestine by the new combination Algorachelus parvus, and in the uppermost Cenomanian of Utah by the new combination Algorachelus tibert. Therefore, a relatively fast and widely distributed geographic dispersion event is recognized, corresponding to the oldest dispersal event so far identified for a lineage of Pleurodira from Gondwana to Laurasia.

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5

Gaffney, Eugene S., G. E. Hooks, and Vincent P. Schneider. "New Material of North American Side-Necked Turtles (Pleurodira: Bothremydidae)." American Museum Novitates 3655 (July 25, 2009): 1–28. http://dx.doi.org/10.1206/626-1.1.

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6

Cadena, Edwin A., Jonathan I. Bloch, and Carlos A. Jaramillo. "New bothremydid turtle (Testudines, Pleurodira) from the Paleocene of northeastern Colombia." Journal of Paleontology 86, no. 4 (July 2012): 688–98. http://dx.doi.org/10.1666/11-128r1.1.

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A new turtle, Puentemys mushaisaensis, from the middle to late Paleocene Cerrejón Formation of Colombia, is described on the basis of a partial skull and many partial to nearly complete carapaces and plastrons representing multiple ontogenetic stages. Whereas P. mushaisaensis is unique in aspects of its shell morphology, it shares many diagnostic characteristics of bothremydid pleurodirans, including a long exoccipital-quadrate contact, a very low and rounded almost circular carapace, and a thinner internal bone cortex than that of the external cortex in both the carapace and plastron. With a maximum carapacial length of 151 cm, P. mushaisaensis is the largest known bothremydid turtle and represents the first occurrence of bothremydids in the Paleogene of South American tropics. Results from a cladistic analysis of bothremydids indicate that P. mushaisaensis shares a close relationship with Foxemys mechinorum from the Late Cretaceous of Europe, indicating a wide-spread geographical distribution for bothremydines during the Late Cretaceous–Paleocene.

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7

GAFFNEY, EUGENE S., HAIYAN TONG, and PETER A. MEYLAN. "EVOLUTION OF THE SIDE-NECKED TURTLES: THE FAMILIES BOTHREMYDIDAE, EURAXEMYDIDAE, AND ARARIPEMYDIDAE." Bulletin of the American Museum of Natural History 300 (December 2006): 1–698. http://dx.doi.org/10.1206/0003-0090(2006)300[1:eotstt]2.0.co;2.

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8

Pérez-García, Adán. "A new member of Taphrosphyini (Pleurodira, Bothremydidae) from the Maastrichtian of Niger." Journal of African Earth Sciences 158 (October 2019): 103548. http://dx.doi.org/10.1016/j.jafrearsci.2019.103548.

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9

Joyce, Walter G., Tyler R. Lyson, and James I. Kirkland. "An early bothremydid (Testudines, Pleurodira) from the Late Cretaceous (Cenomanian) of Utah, North America." PeerJ 4 (September 28, 2016): e2502. http://dx.doi.org/10.7717/peerj.2502.

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BackgroundBothremydidae is a clade of extinct pleurodiran turtles known from the Cretaceous to Paleogene of Africa, Europe, India, Madagascar, and North and South America. The group is most diverse during the Late Cretaceous to Paleogene of Africa. Little is known, however, about the early evolution of the group.MethodsWe here figure and describe a fossil turtle from early Late Cretaceous deposits exposed at MacFarlane Mine in Cedar Canyon, southwestern Utah, USA. The sediments associated with the new turtle are utilized to infer its stratigraphic provenience and the depositional settings in which it was deposited. The fossil is compared to previously described fossil pleurodires, integrated into a modified phylogenetic analysis of pelomedusoid turtles, and the biogeography of bothremydid turtles is reassessed. In light of the novel phylogenetic hypotheses, six previously established taxon names are converted to phylogenetically defined clade names to aid communication.ResultsThe new fossil turtle can be inferred with confidence to have originated from a brackish water facies within the late Cenomanian Culver Coal Zone of the Naturita Formation. The fossil can be distinguished from all other previously described pleurodires and is therefore designated as a new taxon,Paiutemys tibertgen. et. sp. nov. Phylogenetic analysis places the new taxon as sister to the EuropeanPolysternon provinciale,Foxemys trabantiandFoxemys mechinorumat the base of Bothremydinae. Biogeographic analysis suggests that bothremydids originated as continental turtles in Gondwana, but that bothremydines adapted to near-shore marine conditions and therefore should be seen as having a circum-Atlantic distribution.

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10

ROMANO, PEDRO S. R. "The tale of the headless turtle." Zootaxa 4200, no. 2 (November 29, 2016): 327. http://dx.doi.org/10.11646/zootaxa.4200.2.7.

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Pelomedusoides is the most diverse clade of side-necked turtles and there is an extensive fossil record (de Broin, 1988; Lapparent de Broin, 2000; Gaffney et al., 2006, 2011) that dates back at least to the Barremian (Lower Cretaceous) (Romano et al., 2014). Its large fossil record evidences a greater diversity in the past, particularly at the end of the Mesozoic, and exhibits a good sampling of species that are represented by skull material (Gaffney et al., 2006, 2011). As a consequence, the most complete and recent phylogenetic hypotheses for this clade (e.g. Romano et al., 2014; Cadena, 2015) are based on matrices comprising a great amount of cranial characters derived largely from Gaffney et al. (2006, 2011). In addition, it is well established that shell characters show a lot of phenotypic plasticity, even in the fossil species (Romano, 2008; Gaffney et al., 2006, 2011). In most cases it consequently is not justified to rely on “diagnostic features” of poorly informative shell-only material for describing a new species. Because of that, most authors remark new morphotypes in the literature when such aberrant specimens are recovered, but do not make any nomenclatural act by proposing a new yet poorly supported species (e.g. Romano et al., 2013; Ferreira & Langer, 2013; Menegazzo et al., 2015). Unfortunately, such a supposedly new bothremydid turtle (Pleurodira: Bothremydidae) from the Early Paleocene of Brazil was recently described based on poorly diagnostic remains (Carvalho et al., 2016; hereafter CGB, for the authors initials) and a correction of this unfounded nomenclatural act is required. In addition I present some comments on shell only material from Brazil in order to guide splitter-taxonomists to stop describing poorly preserved fossil specimens as new species.

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11

GAFFNEY, EUGENE S., RICHARD T. J. MOODY, and CYRIL A. WALKER. "Azabbaremys, a new Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Paleocene of Mali." American Museum Novitates 3320 (January 2001): 1–16. http://dx.doi.org/10.1206/0003-0082(2001)320<0001:aansnt>2.0.co;2.

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12

Gaffney, Eugene S., Eric Roberts, Famory Sissoko, Mohamed L. Bouaré, Leif Tapanila, and Maureen A. O'leary. "Acleistochelys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Paleocene of Mali." American Museum Novitates 3549, no. 1 (2007): 1. http://dx.doi.org/10.1206/0003-0082(2007)3549[1:aanstp]2.0.co;2.

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13

Pérez-García, Adán, Francisco Ortega, and Xabier Murelaga. "A new genus of Bothremydidae (Chelonii, Pleurodira) in the Cretaceous of Southwestern Europe." Geobios 45, no. 2 (March 2012): 219–29. http://dx.doi.org/10.1016/j.geobios.2011.03.001.

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14

Romano, Pedro S. R., Valéria Gallo, Renato R. C. Ramos, and Luzia Antonioli. "Atolchelys lepida , a new side-necked turtle from the Early Cretaceous of Brazil and the age of crown Pleurodira." Biology Letters 10, no. 7 (July 2014): 20140290. http://dx.doi.org/10.1098/rsbl.2014.0290.

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We report a new pleurodiran turtle from the Barremian Morro do Chaves Formation, Sergipe-Alagoas Basin, Brazil. We tested the phylogenetic position of Atolchelys lepida gen. et sp. nov. by including it in a comprehensive cladistic analysis of pleurodires. The new species is a basal member of Bothremydidae and simultaneously the oldest unambiguous crown Pleurodira. The biogeographic and chronostratigraphic significance of the finding has implications for the calibration of molecular clocks studies by pushing back the minimum age of crown Pleurodira by more than 12 Ma ( ca 125 Ma). The reanalysis of Pelomedusoides relationships provides evidence that the early evolution and relationships among the main lineages of side-necked turtles can be explained, at least partially, by a sequence of vicariance events.

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15

AbdelGawad, Mohamed, Adán Pérez-García, Ren Hirayama, Sara Mohesn, Abdel-Aziz Tantawy, and Gebely Abu El-Kheir. "The First Side-Necked Turtle (Pleurodira, Bothremydidae) from the Campanian (Late Cretaceous) of Egypt." Diversity 15, no. 2 (February 16, 2023): 284. http://dx.doi.org/10.3390/d15020284.

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The Quseir Formation is an Upper Cretaceous (Campanian) deposit in the Kharga oasis of the Southwestern Desert (Egypt). This formation comprises a clastic sequence of bioturbated mudstone and sandstone intercalations, including rare scattered and fragmented vertebrate remains such as shark teeth, dinosaur remains, and turtle plates. These deposits indicate a supratidal marsh environment. A complete shell of a turtle discovered from the Quseir Formation, at the Kharga oasis, is attributed here to Bothremydini (Pleurodira, Bothremydidae), and determined as a new taxon: Khargachelys caironensis gen. et sp. nov. This form represents the only Bothremydini member currently identified in the Campanian record not only of Egypt but also of North Africa. Therefore, it helps fill the missing evolutionary gap from the Late Cretaceous pleurodires in Egypt and in North Africa.

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Gaffney, Eugene S., David W. Krause, and Iyad S. Zalmout. "Kinkonychelys, A New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Madagascar." American Museum Novitates 3662 (August 28, 2009): 1–25. http://dx.doi.org/10.1206/672.1.

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17

GAFFNEY, EUGENE S., DIOGENES DE ALMEIDA CAMPOS, and REN HIRAYAMA. "Cearachelys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Early Cretaceous of Brazil." American Museum Novitates 3319 (January 2001): 1–20. http://dx.doi.org/10.1206/0003-0082(2001)319<0001:cansnt>2.0.co;2.

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18

GAFFNEY, EUGENE S., SANKAR CHATTERJEE, and DHIRAJ K. RUDRA. "Kurmademys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of India." American Museum Novitates 3321 (January 2001): 1–16. http://dx.doi.org/10.1206/0003-0082(2001)321<0001:kansnt>2.0.co;2.

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19

GAFFNEY, EUGENE S., HAIYAN TONG, and PETER A. MEYLAN. "Galianemys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Morocco." American Museum Novitates 3379 (August 2002): 1–20. http://dx.doi.org/10.1206/0003-0082(2002)379<0001:gansnt>2.0.co;2.

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20

GAFFNEY, EUGENE S., ASHOK SAHNI, HERMAN SCHLEICH, SWARN DEEP SINGH, and RAHUL SRIVASTAVA. "Sankuchemys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of India." American Museum Novitates 3405 (May 2003): 1–10. http://dx.doi.org/10.1206/0003-0082(2003)405<0001:sanstp>2.0.co;2.

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21

GAFFNEY, EUGENE S., and CATHERINE A. FORSTER. "Side-Necked Turtle Lower Jaws (Podocnemididae, Bothremydidae) from the Late Cretaceous Maevarano Formation of Madagascar." American Museum Novitates 3397 (March 2003): 1–13. http://dx.doi.org/10.1206/0003-0082(2003)397<0001:stljpb>2.0.co;2.

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22

GAFFNEY, EUGENE S., and CATHERINE A. FORSTER. "Side-Necked Turtle Lower Jaws (Podocnemididae, Bothremydidae) from the Late Cretaceous Maevarano Formation of Madagascar." American Museum Novitates 3397 (March 2003): 1–13. http://dx.doi.org/10.1206/0003-0082(2003)397<1:stljpb>2.0.co;2.

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23

Lehman, Thomas M., and Steven L. Wick. "Chupacabrachelys complexus, n. gen. n. sp. (Testudines: Bothremydidae), from the Aguja Formation (Campanian) of West Texas." Journal of Vertebrate Paleontology 30, no. 6 (December 2, 2010): 1709–25. http://dx.doi.org/10.1080/02724634.2010.520782.

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24

Laurent, Yves, Haiyan Tong, and Julien Claude. "New side-necked turtle (Pleurodira: Bothremydidae) from the Upper Maastrichtian of the Petites-Pyrénées (Haute-Garonne, France)." Cretaceous Research 23, no. 4 (August 2002): 465–71. http://dx.doi.org/10.1006/cres.2002.1015.

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25

Pérez-García, A. "New genera of Taphrosphyina (Pleurodira, Bothremydidae) for the French Maastrichtian ‘Tretosternum’ ambiguum and the Peruvian Ypresian ‘Podocnemis’ olssoni." Historical Biology 32, no. 4 (August 14, 2018): 555–60. http://dx.doi.org/10.1080/08912963.2018.1506779.

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26

RABI, MÁRTON, HAIYAN TONG, and GÁBOR BOTFALVAI. "A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini." Geological Magazine 149, no. 4 (November 4, 2011): 662–74. http://dx.doi.org/10.1017/s0016756811000756.

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AbstractThe continental deposits of the Upper Cretaceous (Santonian) Csehbánya Formation of the Bakony Mountains in Hungary yielded abundant remains of a bothremydid side-necked turtle, which are attributed to a new species of the genus Foxemys, Foxemys trabanti. F. trabanti shows strong affinities with the European monophyletic group Foxemydina owing to the absence of pits in the upper and lower triturating surfaces, the exclusion of the jugal from the triturating surface, the separation of the Eustachian tube and the stapes by a narrow fissure, the presence of deep and narrow fossa pterygoidei, the partially closed foramen jugulare posterius and the pentagonal shape of the basisphenoid in ventral view. Among the Foxemydina the bothremydid from Iharkút is more closely related to F. mechinorum than to Polysternon provinciale from the Early Campanian of France, mainly because of the position of the occipital condyle relative to the mandibular condyles of the quadrate. The new remains represent the only record of the Foxemydina outside of Western Europe and provide the earliest known occurrence of this endemic, freshwater group in the former Mediterranean Basin. The historical biogeography of the tribe Bothremydini is investigated and a hypothesis of migration from Africa to North America via the high-latitude Thulean route is put forward.

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27

García, Adán Pérez, Francisco Ortega, and Xabier Murelaga. "Ampliación de la distribución geográfica y temporal deElochelys convenarum(Chelonii, Bothremydidae) en el Cretácico Superior de la Península Ibérica." Ameghiniana 47, no. 3 (September 2010): 307–15. http://dx.doi.org/10.5710/amgh.v47i3.6.

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28

CARVALHO, ANNY RAFAELA DE ARAÚJO, ALINE MARCELE GHILARDI, and ALCINA MAGNÓLIA FRANCA BARRETO. "A new side-neck turtle (Pelomedusoides: Bothremydidae) from the Early Paleocene (Danian) Maria Farinha Formation, Paraíba Basin, Brazil." Zootaxa 4126, no. 4 (June 21, 2016): 491. http://dx.doi.org/10.11646/zootaxa.4126.4.3.

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29

Pérez-García, Adán. "A new turtle taxon (Podocnemidoidea, Bothremydidae) reveals the oldest known dispersal event of the crown Pleurodira from Gondwana to Laurasia." Journal of Systematic Palaeontology 15, no. 9 (September 28, 2016): 709–31. http://dx.doi.org/10.1080/14772019.2016.1228549.

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30

Raveloson, Miky Lova Tantely. "Two Allopatric Bothremydidae Taxa of Turtle in the India-Madagascar Faunal Provinces During the Late Cretaceous: Evidence of Biotic Dispersal Across the Microcontinent." Paleontological Society Special Publications 13 (2014): 20–21. http://dx.doi.org/10.1017/s247526220001039x.

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31

Maniel, I. J., M. S. de la Fuente, and J. I. Canale. "The first Cearachelyini (Pelomedusoides, Bothremydidae) turtle from the Upper Cretaceous of Patagonia, and an overview of the occurrence and diversity of Pelomedusoides in Patagonia." Cretaceous Research 125 (September 2021): 104869. http://dx.doi.org/10.1016/j.cretres.2021.104869.

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32

Bardet, Nathalie. "Maastrichtian marine reptiles of the Mediterranean Tethys: a palaeobiogeographical approach." Bulletin de la Société Géologique de France 183, no. 6 (December 1, 2012): 573–96. http://dx.doi.org/10.2113/gssgfbull.183.6.573.

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AbstractA global comparison of coeval Maastrichtian marine reptiles (squamates, plesiosaurs, chelonians and crocodyliformes) of Europe, New Jersey, northwestern Africa and Middle-East has been performed. More than twenty outcrops and fifty species (half of them being mosasaurids) have been recorded. PEA and Cluster Analysis have been performed using part of this database and have revealed that marine reptile faunas (especially the mosasaurid ones) from the Mediterranean Tethys are clearly segregated into two different palaeobiogeographical provinces: 1) The northern Tethys margin province (New Jersey and Europe), located around palaeolatitudes 30-40°N and developping into warm-temperate environments, is dominated by mosasaurid squamates and chelonioid chelonians; it is characterized by the mosasaurid association of Mosasaurus hoffmanni and Prognathodon sectorius. 2) The southern Tethys margin province (Brazil and the Arabo-African domain), located between palaeolatitudes 20°N-20°S and developping into intertropical environments, is dominated by mosasaurid squamates and bothremydid chelonians; it is characterized by the mosasaurid association of Globidens phosphaticus as well as by Halisaurus arambourgi and Platecarpus (?) ptychodon (Arabo-African domain). These faunal differences are interpreted as revealing palaeoecological preferences probably linked to differences in palaeolatitudinal gradients and/or to palaeocurrents.On a palaeoecological point on view and concerning mosasaurids, the mosasaurines (Prognathodon, Mosasaurus, Globidens and Carinodens) prevail on both margins but with different species. The ichthyophageous plioplatecarpines Plioplatecarpus (Northern margin) and Platecarpus (?) ptychodon (Southern margin) characterise respectively each margin. The halisaurine Halisaurus is present on both margins but with different species. Of importance, the tylosaurines remain currently unknown on the southern Tethys margin and are restricted to higher palaeolatitudes. Chelonians (bothremydids and chelonioids) are respective of each margin, which probably indicates lower dispersal capabilities compared to mosasaurids. The relative scarcity of plesiosaurs and crocodyliformes could be linked to different ecological preferences. The noteworthy crocodyliforme diversity increase in the Palaeogene is probably linked to mosasaurid extinction during the biological crisis of the K/Pg boundary.

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KEAR, B. P., T. H. RICH, M. A. ALI, Y. A. AL-MUFARRIH, A. H. MATIRI, A. M. MASARY, and Y. ATTIA. "Late Cretaceous (Campanian—Maastrichtian) marine reptiles from the Adaffa Formation, NW Saudi Arabia." Geological Magazine 145, no. 5 (June 11, 2008): 648–54. http://dx.doi.org/10.1017/s0016756808005062.

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AbstractMarine reptile remains occur in the Upper Cretaceous (lower Campanian to lower Maastrichtian) Adaffa Formation of NW Saudi Arabia. This is the first detailed report of late Mesozoic marine reptiles from the Arabian Peninsula. The fossils include bothremydid (cf. Taphrosphyini) turtles, dyrosaurid crocodyliforms, elasmosaurid plesiosaurs, mosasaurs (Prognathodon, plioplatecarpines) and an indeterminate small varanoid. The assemblage is compositionally similar to contemporary faunas from elsewhere in the Middle East/North Africa, and comprises taxa that are typical of the southern margin of the Mediterranean Tethys.

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Pérez-García, A. "A new bothremydid turtle (Pleurodira) from the Upper Cretaceous (Cenomanian) of Madagascar." Cretaceous Research 118 (February 2021): 104645. http://dx.doi.org/10.1016/j.cretres.2020.104645.

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KEAR, BENJAMIN P., and GRAHAM E. BUDD. "New perspectives on ancient marine reptiles." Geological Magazine 151, no. 1 (October 25, 2013): 5–6. http://dx.doi.org/10.1017/s0016756813000873.

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Amniotes first invaded saline lagoons and coastal seaways towards the end of the Palaeozoic (Early Permian, ~ 280 Ma: Piñeiro et al. 2012), but by the dawn of the Mesozoic (Early–Middle Triassic, ~ 250–235 Ma: Rieppel, 2002; McGowan & Motani, 2003) they had achieved a diversity of specialized body-forms requisite for an obligate oceanic lifestyle. Such an explosive ecomorphological radiation paved the way for amniote dominance of large-bodied aquatic carnivore/omnivore niches over the next 185 Ma, with some lineages (e.g. dyrosaurid crocodylomorphs and bothremydid turtles: Gaffney, Tong & Meylan, 2006; Barbosa, Kellner & Sales Viana, 2008) even persisting on into the Palaeogene (until ~ 50 Ma), and diversifying (i.e. chelonioid sea turtles: Hirayama, 1997) alongside emergent marine mammals through the Neogene (from ~ 23 Ma) and up until today.

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Pérez-García, Adán, and Francisco Ortega. "Identification of the French Upper Cretaceous bothremydid turtle Foxemys mechinorum in the Spanish record." Geobios 51, no. 3 (June 2018): 211–17. http://dx.doi.org/10.1016/j.geobios.2018.04.002.

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López-Conde, Oliver A., Adán Pérez-García, María L. Chavarría-Arellano, and Jesús Alvarado-Ortega. "A new bothremydid turtle (Pleurodira) from the Olmos Formation (upper Campanian) of Coahuila, Mexico." Cretaceous Research 119 (March 2021): 104710. http://dx.doi.org/10.1016/j.cretres.2020.104710.

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Pérez García, Adán, Florias Mees, and Thierry Smith. "Shell anatomy of the African Paleocene bothremydid turtle Taphrosphys congolensis and systematic implications within Taphrosphyini." Historical Biology 32, no. 3 (July 18, 2018): 376–85. http://dx.doi.org/10.1080/08912963.2018.1497023.

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Botfalvai, Gábor, Edina Prondvai, and Attila Ősi. "Inferred bite marks on a Late Cretaceous (Santonian) bothremydid turtle and a hylaeochampsid crocodilian from Hungary." Cretaceous Research 50 (July 2014): 304–17. http://dx.doi.org/10.1016/j.cretres.2014.05.006.

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Pérez-García, Adán. "First evidence of a bothremydid turtle (crown Pleurodira) in the middle Cretaceous of Castile and Leon (Spain)." Journal of Iberian Geology 46, no. 4 (November 27, 2020): 363–68. http://dx.doi.org/10.1007/s41513-020-00146-9.

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Marmi, J., Á. H. Luján, V. Riera, R. Gaete, O. Oms, and À. Galobart. "The youngest species of Polysternon: A new bothremydid turtle from the uppermost Maastrichtian of the southern Pyrenees." Cretaceous Research 35 (June 2012): 133–42. http://dx.doi.org/10.1016/j.cretres.2011.12.004.

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Pérez-García, A., M. T. Antunes, F. Barroso-Barcenilla, P. M. Callapez, M. Segura, A. F. Soares, and A. Torices. "A bothremydid from the middle Cenomanian of Portugal identified as one of the oldest pleurodiran turtles in Laurasia." Cretaceous Research 78 (October 2017): 61–70. http://dx.doi.org/10.1016/j.cretres.2017.05.031.

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BARDET, N., H. CAPPETTA, X. PEREDA SUBERBIOLA, M. MOUTY, A. K. AL MALEH, A. M. AHMAD, O. KHRATA, and N. GANNOUM. "The marine vertebrate faunas from the Late Cretaceous phosphates of Syria." Geological Magazine 137, no. 3 (May 2000): 269–90. http://dx.doi.org/10.1017/s0016756800003988.

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Marine vertebrate faunas from the latest Cretaceous phosphates of the Palmyrides Chain of Syria are described for the first time. Recent fieldwork in the phosphatic deposits of the Palmyra area (mines of Charquieh and Khneifiss, outcrops of Bardeh, Soukkari and Soukhneh) have yielded a rich and diversified assemblage of marine vertebrates, including more than 50 species of chondrichthyes, osteichthyes, squamates, chelonians, plesiosaurians and crocodilians. Selachians are the most abundant and diverse component of the faunas and are represented by at least 34 species of both sharks and rays. Actinopterygians include representatives of six families, the most common being the enchodontids. Squamates are known by six mosasaurid species and an indeterminate varanoid. Chelonians are represented by at least two bothremydids and two chelonioids. Finally, elasmosaurid plesiosaurs and indeterminate crocodilians are also present in the fossil assemblages. The difference in faunal composition observed between the sites is interpreted as being due to palaeoecological preferences related to the Hamad Uplift palaeostructure. The marine vertebrate faunas of Syria show close affinities with those of the latest Cretaceous phosphatic deposits of North Africa and the Middle East and are typical of the southern Tethyan realm. From a biostratigraphical point of view, the selachians are the only suitable material to provide elements of an answer to the long debated question of the age of the Syrian Senonian phosphates. They suggest an Early Maastrichtian age for most of the phosphates of the Palmyrides Chain.

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Gaffney, Eugene S., and Haiyan Tong. "Redescription of the Skull of Ummulisani rutgersensis Gaffney, Tong, and Meylan, 2006, a Bothremydid Side-Necked Turtle from the Eocene of Morocco." American Museum Novitates 3615, no. 1 (2008): 1. http://dx.doi.org/10.1206/607.1.

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Joyce, Walter G., and Saswati Bandyopadhyay. "A revision of the pelomedusoid turtle Jainemys pisdurensis from the Late Cretaceous (Maastrichtian) Lameta Formation of India." PeerJ 8 (June 22, 2020): e9330. http://dx.doi.org/10.7717/peerj.9330.

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Background Jainemys pisdurensis comb. nov. is an extinct pleurodiran turtle from the Late Cretaceous (Maastrichtian) of India, previously referred to Carteremys and Shweboemys. The holotype, an eroded skull, had been collected near the village of Pisdura, south of Nagpur, in Maharashtra State, while all referred shell material originates from coeval sediments exposed at the nearby village of Dongargaon. Initial estimates believed this turtle to either be an early representative of Podocnemididae or a basal representative of Pelomedusoides. Methods We here figure and describe all specimens that had previously been referred to Jainemys pisdurensis comb. nov. We furthermore re-evaluate the validity of this fossil turtle and explore its phylogenetic relationships within Pleurodira. Results The holotype of Jainemys pisdurensis comb. nov. displays a morphology that differs substantially from that originally reported. Most notably, the palatines only have a minor contribution to the broad triturating surfaces but have a broad midline contact with each other, the pterygoids only have a midline contact of intermediate length and do not contact the opisthotics posteriorly, the basisphenoid is broad and short, and the opisthotics do not contribute to the flooring of the cavum acustico-jugulare. The referred shell material also displays a morphology different from that reported originally, in particular in that vertebral I does not contribute to the anterior margin of the carapace while the nuchal does. Phylogenetic analysis places the cranial material within the bothremydid clade Kurmademydini, while the shell material is placed in an unresolved polytomy at the base of this clade. Jainemys pisdurensis is confirmed to be a valid species of pleurodiran turtle, but the high diversity of coeval kurmademydines in India demands removal of the postcranial remains from this taxon. The realization that all valid species of Late Cretaceous (Maastrichtian) turtles from India form a clade supports the hypothesis that India was physically separated from the rest of Gondwana at this time.

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PÉREZ-GARCÍA, Adán, Francisco ORTEGA, and Xabier MURELAGA. "Iberoccitanemys atlanticum (Lapparent de Broin & Murelaga, 1996) n. comb.: new data on the diversity and paleobiogeographic distributions of the Campanian-Maastrichtian bothremydid turtles of Europe." Comptes Rendus Palevol, no. 32 (August 16, 2021). http://dx.doi.org/10.5852/cr-palevol2021v20a32.

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Bothremydidae is the most abundant clade of turtles in the Campanian and Maastrichtian (Upper Cretaceous) fossil record of southwestern Europe. Several members of Foxemydina Gaffney, Tong & Meylan, 2006 are known in an area that includes Southern France and the North-Eastern half of Spain. The problematic ‘Polysternon’ atlanticum is the worst characterized, lacking a diagnosis that allows its specific validity to be confirmed, and whose generic attribution has been recognized as doubtful. Its presence was exclusively proposed in its type locality, the upper Campanian quarry of Laño, in Treviño County (Burgos Province, North of Spain). Despite the fact that knowledge about Bothremydidae has markedly increased after the description of ‘Polysternon’ atlanticum Lapparent de Broin & Murelaga, 1996, no new information about this species has been published since the 1990s. The analysis of abundant unpublished material of the bothremydid from Laño allows us to confirm the validity of this species. As a consequence of this study, it is not only identified in its type locality, but also in other Spanish regions and in the south of France. The diversity of Bothremydidae Baur, 1891 in the Upper Cretaceous of Europe is lower than previously considered. Thus, the species ‘Iberoccitanemys convenarum’ (Laurent, Tong & Claude, 2002), originally defined for the French record, and subsequently also identified in Spain, is identified here as a synonym of the species described in Laño. An emended diagnosis for the upper Campanian to upper Maastrichtian, Iberoccitanemys atlanticum (Lapparent de Broin & Murelaga, 1996) n. comb., is proposed.

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Adrian, Brent, Heather F. Smith, Christopher R. Noto, and Aryeh Grossman. "An early bothremydid from the Arlington Archosaur Site of Texas." Scientific Reports 11, no. 1 (May 20, 2021). http://dx.doi.org/10.1038/s41598-021-88905-1.

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AbstractFour turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of the Lewisville Formation (Woodbine Group) in Texas. Herein, we describe a new side-necked turtle (Pleurodira), Pleurochayah appalachius gen. et sp. nov., which is a basal member of the Bothremydidae. Pleurochayah appalachius gen. et sp. nov. shares synapomorphic characters with other bothremydids, including shared traits with Kurmademydini and Cearachelyini, but has a unique combination of skull and shell traits. The new taxon is significant because it is the oldest crown pleurodiran turtle from North America and Laurasia, predating bothremynines Algorachelus peregrinus and Paiutemys tibert from Europe and North America respectively. This discovery also documents the oldest evidence of dispersal of crown Pleurodira from Gondwana to Laurasia. Pleurochayah appalachius gen. et sp. nov. is compared to previously described fossil pleurodires, placed in a modified phylogenetic analysis of pelomedusoid turtles, and discussed in the context of pleurodiran distribution in the mid-Cretaceous. Its unique combination of characters demonstrates marine adaptation and dispersal capability among basal bothremydids.

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Antunes, Miguel Telles, and Adán Pérez‐García. "A bothremydid turtle (Pleurodira) from the middle Cenomanian of Vale de Figueira (Belas, Portugal)." Anatomical Record, March 18, 2024. http://dx.doi.org/10.1002/ar.25429.

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AbstractAn unpublished turtle shell from the middle Cenomanian of Vale de Figueira, near Belas (Lisbon District, Portugal), is recognized by us as collected in 1880 under the direction of Carlos Ribeiro. No turtle remains from that region had so far been figured, described or discussed from a systematic point of view. The specimen corresponds to a partial but articulated shell. It is attributed to Pleurodira and, more specifically, to Bothremydidae. Only one pre‐Campanian turtle specimen was previously identified at the infrafamilial level in the Cretaceous record of Portugal. It was a partial shell from the middle Cenomanian of Nazaré (Leiria District) attributed to the bothremydid Algorachelus peregrina, a species defined in a Spanish synchronous locality (Algora, in Central Spain). Several anatomical regions in the specimen from Vale de Figueira were not preserved in the partial carapace from Nazaré, and differences in the morphology of some elements are recognized between both shells. However, the individual studied here is also ascribed to Algorachelus peregrina, these differences being justified by intraspecific variability. Therefore, the specimen represents the second evidence on the species in Portugal, being the only one recognized for the Lisbon District.

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Joyce, Walter G., Yann Rollot, Serjoscha W. Evers, Tyler R. Lyson, Lydia J. Rahantarisoa, and David W. Krause. "A new pelomedusoid turtle, Sahonachelys mailakavava , from the Late Cretaceous of Madagascar provides evidence for convergent evolution of specialized suction feeding among pleurodires." Royal Society Open Science 8, no. 5 (May 2021). http://dx.doi.org/10.1098/rsos.210098.

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The Maevarano Formation in northwestern Madagascar has yielded a series of exceptional fossils over the course of the last three decades that provide important insights into the evolution of insular ecosystems during the latest Cretaceous (Maastrichtian). We here describe a new genus and species of pelomedusoid turtle from this formation, Sahonachelys mailakavava , based on a nearly complete skeleton. A phylogenetic analysis suggests close affinities of Sahonachelys mailakavava with the coeval Madagascan Sokatra antitra . These two taxa are the only known representatives of the newly recognized clade Sahonachelyidae , which is sister to the speciose clade formed by Bothremydidae and Podocnemidoidae . A close relationship with coeval Indian turtles of the clade Kurmademydini is notably absent. A functional assessment suggests that Sahonachelys mailakavava was a specialized suction feeder that preyed upon small-bodied invertebrates and vertebrates. This is a unique feeding strategy among crown pelomedusoids that is convergent upon that documented in numerous other clades of turtles and that highlights the distinct evolutionary pathways taken by Madagascan vertebrates.

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Pérez-García, Adán. "A new turtle confirms the presence of Bothremydidae (Pleurodira) in the Cenozoic of Europe and expands the biostratigraphic range of Foxemydina." Science of Nature 103, no. 7-8 (June 4, 2016). http://dx.doi.org/10.1007/s00114-016-1375-y.

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